Conophytum Piluliforme 1
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''Conophytum'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of South African and Namibian
succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
plants that belong to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Aizoaceae The Aizoaceae (), or fig-marigold family, is a large family of dicotyledonous flowering plants containing 135 genera and about 1,800 species. Several genera are commonly known as 'ice plants' or 'carpet weeds'. The Aizoaceae are also referred to a ...
. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in
Afrikaans Afrikaans is a West Germanic languages, West Germanic language spoken in South Africa, Namibia and to a lesser extent Botswana, Zambia, Zimbabwe and also Argentina where there is a group in Sarmiento, Chubut, Sarmiento that speaks the Pat ...
), waterblasies (water blisters in Afrikaans), sphaeroids, conos, cone plants, dumplings, or button plants.


Description

''Conophytum'' species are dwarf cushion-forming or single-bodied succulents. Members of the genus are tiny plants with succulent leaves ranging from 1/4" to 2" in length. These leaves are partially or entirely fused along their centers. Each leaf pair (together referred to as a body) ranges in shape from "bilobed" to spherical to ovoid to tubular to conical. Some species have
epidermal window Leaf window, also known as epidermal window, and fenestration, is a specialized leaf structure consisting of a translucent area through which light can enter the interior surfaces of the leaf where photosynthesis can occur. The translucent struct ...
s on the top of their leaves. To the naked eye the epidermis ranges from very smooth to slightly rough to hairy, depending on the microscopic epidermal cell shape and structure. In their normal, natural state each stem has only one pair of leaves at a time though one plant may have dozens of stems and thus dozens of leaf pairs. When very heavy rains come to their native habitat they may grow luxuriantly and develop two leaf pairs per stem simultaneously; this is called "stacking up" of the leaves. The plants grow very slowly; depending on the species, a 50-year-old plant might not be bigger than a
walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...
.


Taxonomy

The genus is sometimes wrongly referred to as ''Conophyton'', the name that
Adrian Hardy Haworth Adrian Hardy Haworth (19 April 1767, in Kingston upon Hull, Hull – 24 August 1833, in Chelsea, London, Chelsea) was an England, English entomologist, botanist and carcinologist. Family The younger son of Benjamin Haworth, of Haworth Hall and ...
suggested in 1821: "If this section proves to be a genus, the name of Conophyton would be apt". However, this was too tentative to establish a validly published generic name and also, Haworth himself neither adopted it nor accepted the genus. The genus was neither recognised nor validly named until the name ''Conophytum'' was published 101 years later. It was published by
Nicholas Edward Brown Nicholas Edward Brown (11 July 1849 in Redhill, Surrey – 25 November 1934 in Kew Gardens, London) was an English plant taxonomist and authority on succulents. He was also an authority on several families of plants, including Asclepiadaceae, ...
in 1922.''Conophytum'' N.E.Br. (n.d.). International Plant Names Index. Retrieved January 27, 2025, from https://www.ipni.org/n/16140-1 The lectotype ''
Conophytum minutum ''Conophytum minutum'', called the lesser dumpling, is a species of flowering plant in the genus ''Conophytum'', native to the western Cape Provinces of South Africa. It has gained the Royal Horticultural Society's Award of Garden Merit The Aw ...
'' was designated in 1951.


Species

This is a list of species in the genus ''Conophytum''. *''
Conophytum achabense ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum acutum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum albiflorum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum angelicae ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum armianum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family (biology), family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as kno ...
'' *''
Conophytum arthurolfago ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum auriflorum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum bachelorum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum bicarinatum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum bilobum ''Conophytum bilobum'' is a plant in the family Aizoaceae The Aizoaceae (), or fig-marigold family, is a large family of dicotyledonous flowering plants containing 135 genera and about 1,800 species. Several genera are commonly known as 'ice pl ...
'' *''
Conophytum blandum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum bolusiae ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum breve ''Conophytum breve'' is a small South African species of succulent plant of the genus ''Conophytum''. Description ''Conophytum breve'' has small, smooth, rounded heads, and offsets to form irregular clumps. The epidermis is a chalky grey to glau ...
'' *''
Conophytum brunneum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum bruynsii ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *''
Conophytum burgeri ''Conophytum burgeri'' ("Burger's onion") is a small, endangered, South African species of succulent plant, of the genus '' Conophytum''. Description An onion-shaped, single-bodied, succulent plant, it is possibly the most unusual of all the s ...
'' *'' Conophytum calculus'' *'' Conophytum caroli'' *''
Conophytum carpianum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family (biology), family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as kno ...
'' *'' Conophytum chauviniae'' *'' Conophytum chrisocruxum'' *''
Conophytum chrisolum ''Conophytum chrisolum'' is a species of succulent plant in the ''Iceplant Family'' Aizoaceae. It is endemic to the Richtersveld region of Northern Cape Province in South Africa South Africa, officially the Republic of South Africa (RSA ...
'' *'' Conophytum comptonii'' *'' Conophytum concavum'' *'' Conophytum concordans'' *''
Conophytum cubicum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum depressum'' *''
Conophytum devium ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum ectypum'' *'' Conophytum ernstii'' *'' Conophytum ficiforme'' *'' Conophytum flavum'' *'' Conophytum francoiseae'' *'' Conophytum fraternum'' *'' Conophytum friedrichiae'' *'' Conophytum frutescens'' *'' Conophytum fulleri'' *'' Conophytum globosum'' *'' Conophytum halenbergense'' *'' Conophytum hammeri'' *''
Conophytum hermarium ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum herreanthus'' *'' Conophytum hians'' *'' Conophytum irmae'' *'' Conophytum jarmilae'' *'' Conophytum joubertii'' *'' Conophytum jucundum'' *'' Conophytum khamiesbergense'' *'' Conophytum klinghardtense'' *'' Conophytum kubusanum'' *'' Conophytum limpidum'' *'' Conophytum lithopsoides'' *'' Conophytum loeschianum'' *'' Conophytum longibracteatum'' *''
Conophytum longum ''Conophytum longum'', called the long-leaved cone plant, is a species of flowering plant in the genus ''Conophytum'', native to the western Cape Provinces of South Africa. It has gained the Royal Horticultural Society's Award of Garden Merit. R ...
'' *'' Conophytum luckhoffii'' *'' Conophytum lydiae'' *'' Conophytum marginatum'' *'' Conophytum maughanii'' *'' Conophytum meyeri'' *'' Conophytum minimum'' *'' Conophytum minusculum'' *''
Conophytum minutum ''Conophytum minutum'', called the lesser dumpling, is a species of flowering plant in the genus ''Conophytum'', native to the western Cape Provinces of South Africa. It has gained the Royal Horticultural Society's Award of Garden Merit The Aw ...
'' *'' Conophytum mirabile'' *''
Conophytum obcordellum ''Conophytum obcordellum'' is a species of flowering plant in the family Aizoaceae, native to the Western Cape of South Africa. It is a small clump-forming succulent, growing to 10 cm tall and 1 m broad, forming glossy grey pebble-shaped ...
'' *''
Conophytum obscurum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum pageae'' *'' Conophytum pellucidum'' *'' Conophytum phoeniceum'' *'' Conophytum piluliforme'' *'' Conophytum praesectum'' *'' Conophytum pubescens'' *'' Conophytum pubicalyx'' *'' Conophytum quaesitum'' *'' Conophytum ratum'' *'' Conophytum reconditum'' *'' Conophytum regale'' *'' Conophytum ricardianum'' *'' Conophytum roodiae'' *'' Conophytum rugosum'' *'' Conophytum saxetanum'' *'' Conophytum schlechteri'' *'' Conophytum semivestitum'' *'' Conophytum smorenskaduense'' *'' Conophytum stephanii'' *'' Conophytum stevens-jonesianum'' *'' Conophytum subfenestratum'' *'' Conophytum subterraneum'' *'' Conophytum swanepoelianum'' *''
Conophytum tantillum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum taylorianum'' *'' Conophytum tomasi'' *'' Conophytum truncatum'' *''
Conophytum turrigerum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
'' *'' Conophytum uviforme'' *'' Conophytum vanheerdei'' *'' Conophytum velutinum'' *'' Conophytum verrucosum'' *'' Conophytum violaciflorum'' *'' Conophytum wettsteinii''


Conservation

Recently several species have been threatened with extinction due to mining and poaching from the wild for the black market. These plants are mainly sold to collectors in Asian countries, where there has been a high demand for them.Oder, T. (2018, December 1). Why the Black Market for Cacti and Succulents Is Booming. Treehugger. Retrieved January 27, 2025, from https://www.treehugger.com/why-black-market-cacti-succulents-booming-4863289 In June 2021, police arrested a dozen men at a farm in the Western Cape for possession of 4,000 ''
Conophytum acutum ''Conophytum'' is a genus of South African and Namibian succulent plants that belong to the family Aizoaceae. The name is derived from the Latin ''conus'' (cone) and Greek ''phytum'' (plant). The plants are also known as knopies (buttons in A ...
''; this came as a surprise as scientists thought there were only about a thousand of the plants in the wild and none on the farm in question. Teams at the Kirstenbosch Botanical Garden are collecting and categorising seeds from ''Conophytum'' and other succulents in partnership with the
Millennium Seed Bank The Millennium Seed Bank Partnership (MSBP or MSB), formerly known as the Millennium Seed Bank Project, is the largest '' ex situ'' plant conservation programme in the world, coordinated by the Royal Botanic Gardens, Kew, UK. After being awar ...
.


Cultivation

Several species of ''Conophytum'' are found in cultivation. ''Conophytum'' grows best between the temperatures and . When the plant is not under optimal temperatures, its leaves will curl and lose color. If grown in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, the plants should not be watered from the end of March to late July. During this time, leaves will dry and protect the newer leaves. It is recommended to water ''Conophytum'' every week from late July/early August until mid October, where watering should be reduced to once every three weeks throughout winter. Weekly watering can resume when the temperature increases in February.


References

{{Authority control Aizoaceae genera Flora of Namibia Flora of South Africa Flora of the Cape Provinces Succulent plants Taxa named by N. E. Brown