The common torpedo (''Torpedo torpedo''), also known as ocellate torpedo or eyed electric ray, is a

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

electric ray The electric rays are a group of rays, flattened cartilaginous fish with enlarged pectoral fins, composing the order Torpediniformes . They are known for being capable of producing an electric discharge, ranging from 8 to 220 volts, depending ...

in the family Torpedinidae. It is found in the

Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ...

and the eastern

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth#Surface, Earth's surface and about 29% of its water surface area. It is known to separate the "Old World" of Afr ...

from the

Bay of Biscay The Bay of Biscay (), known in Spain as the Gulf of Biscay ( es, Golfo de Vizcaya, eu, Bizkaiko Golkoa), and in France and some border regions as the Gulf of Gascony (french: Golfe de Gascogne, oc, Golf de Gasconha, br, Pleg-mor Gwaskogn), ...

Angola , national_anthem = "Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...

, and is a benthic fish typically encountered over soft substrates in fairly shallow, coastal waters. Growing to long, this species has a nearly circular

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

disc and a short, thick tail with two

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

s of nearly equal size and a large

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...

. It can be identified by the prominent blue spots on its back, which usually number five but may vary from zero to nine, as well as by the small knobs on the rims of its spiracles. For attack and defense, the common torpedo can deliver a strong

electric shock Electrical injury is a physiological reaction caused by electric current passing through the body. The injury depends on the density of the current, tissue resistance and duration of contact. Very small currents may be imperceptible or produce a ...

of up to 200

volt The volt (symbol: V) is the unit of electric potential, electric potential difference ( voltage), and electromotive force in the International System of Units (SI). It is named after the Italian physicist Alessandro Volta (1745–1827). D ...

s. It is a solitary,

nocturnal Nocturnality is an ethology, animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatures generally have ...

ambush predator Ambush predators or sit-and-wait predators are carnivorous animals that capture or trap prey via stealth, luring or by (typically instinctive) strategies utilizing an element of surprise. Unlike pursuit predators, who chase to capture prey u ...

that feeds mainly on

bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartila ...

es and

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...

s. This species is

aplacental viviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

, with the developing

embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm ...

s nourished by

yolk Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example bec ...

and histotroph ("uterine milk") produced by the mother. Females produce litters of up to 28 young every year in late summer or autumn, after a

gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once ...

of 4–8 months; the specifics vary between geographical regions. The common torpedo's shock is painful but otherwise of little danger to humans. Its electrogenic properties led it to be used in

medicine Medicine is the science and Praxis (process), practice of caring for a patient, managing the diagnosis, prognosis, Preventive medicine, prevention, therapy, treatment, Palliative care, palliation of their injury or disease, and Health promotion ...

by the

ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...

s and Romans. In modern times, it has no economic value and is mostly discarded when caught as

bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juven ...

in fisheries. The impact of fishing on its population is uncertain, and the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natur ...

(IUCN) has listed it as Vulnerable.

Taxonomy

The common torpedo and other electric rays were familiar to the peoples of

classical antiquity Classical antiquity (also the classical era, classical period or classical age) is the period of cultural history between the 8th century BC and the 5th century AD centred on the Mediterranean Sea, comprising the interlocking civilizations ...

. ''Torpedo'' was the

Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD * Roman people, the people of ancient Rome *'' Epistle to the Romans'', shortened to ''Romans'', a lett ...

name for electric rays, derived from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...

meaning 'to be numb'.

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...

, known as the "father of

taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

", scientifically described the common torpedo as ''Raja torpedo'' in the 1758 tenth edition of his ''

Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nom ...

''. However, the common torpedo also appeared in at least 52 pre-Linnaean sources under various names such as ''Torpedo'', ''Raja tota lævis'', ''Torpedo maculosa'', and ''Torpedo Sinûs Persici''. These early accounts, including Linnaeus's, confounded the common torpedo with other electric ray species. As Linnaeus did not indicate any

type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

s, the designation of a

lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...

neotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...

is warranted in the interest of taxonomic stability. This measure has yet to be taken.

André Marie Constant Duméril André Marie Constant Duméril (1 January 1774 – 14 August 1860) was a French zoologist. He was professor of anatomy at the Muséum national d'histoire naturelle from 1801 to 1812, when he became professor of herpetology and ichthyology. ...

was the first author to refer to ''Torpedo'' as a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

, in his 1806 ''Zoologie analytique, ou méthode naturelle de classification des animaux''. Duméril did not name any ''Torpedo'' species; the first author to do so was probably

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...

, who assigned Linnaeus's ''Raja torpedo'' to the genus ''Torpedo'' in 1838. Since at the time ''T. torpedo'' was the only member of the genus, it became the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

. Within the genus ''Torpedo'', the common torpedo is placed within the subgenus ''Torpedo'', which differs from the other subgenus ''Tetronarce'' in having spiracles with

papillate This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...

rims and ornate dorsal coloration. This species may also be referred colloquially to as crampfish, cramp ray, or torpedo ray.

Distribution and habitat

Native to the eastern

, the common torpedo occurs from the

Bay of Biscay The Bay