Common Tody-flycatcher
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The common tody-flycatcher (''Todirostrum cinereum'') is a small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the family
Tyrannidae The tyrant flycatchers (Tyrannidae) comprise a Family (biology), family of passerine birds which is found virtually throughout North America, North and South America. It is the world's largest family of birds, with more than 400 species, and is ...
, the tyrant flycatchers. It is found in
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, in every
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
n country, and in every mainland
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
n country except
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
and
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
.


Taxonomy and systematics

The common tody-flycatcher was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1766 in the
twelfth edition Twelfth can mean: *The Twelfth Amendment to the United States Constitution *The Twelfth, a Protestant celebration originating in Ireland In mathematics: * 12th, an ordinal number; as in the item in an order twelve places from the beginning, follo ...
of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Todus cinereus''. Linnaeus based his description on the "Grey and Yellow Fly-catcher" that had been described and illustrated in 1760 by George Edwards from a specimen collected in
Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...
. The specific epithet ''cinereum'' is from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''cinereus'' meaning "ash-grey". The common tody-flycatcher is now the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of the genus ''
Todirostrum ''Todirostrum'' is a genus of Neotropical birds in the New World flycatcher family (biology), family Tyrannidae. Taxonomy and species list The genus ''Todirostrum'' was erected in 1831 by the French naturalist René Lesson. The type species was ...
'' that was introduced by
René Lesson René Primevère Lesson (20 March 1794 – 28 April 1849) was a French surgery, surgeon, natural history, naturalist, ornithologist, and herpetologist. Biography Lesson was born at Rochefort, Charente-Maritime, Rochefort, and entered the Naval ...
in 1831. Eight
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognized: * ''T. c. virididorsale''
Parkes Parkes may refer to: * Sir Henry Parkes (1815–1896), Australian politician, one of the earliest and most prominent advocates for Australian federation Named for Henry Parkes * Parkes, New South Wales, a regional town * Parkes Observatory, a radi ...
, 1976
* ''T. c. finitimum''
Bangs Bang, bang!, or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Central African ...
, 1904
* ''T. c. wetmorei'' Parkes, 1976 * ''T. c. sclateri'' (
Cabanis Cabanis is a surname. Notable people with the surname include: * Bernard Cabanis (born 1950), French ice hockey player * George Cabanis (1815–1892), American politician * James Cabanis (1838–1920), American politician, son of George Cabanis *Jea ...
&
Heine Heine is both a surname and a given name of German origin. People with that name include: People with the surname * Albert Heine (1867–1949), German actor * Alice Heine (1858–1925), American-born princess of Monaco * Armand Heine (1818–1883) ...
, 1860)
* ''T. c. cinereum'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1766)
* ''T. c. peruanum'' Zimmer, JT, 1930 * ''T. c. coloreum'' Ridgway, 1906 * ''T. c. cearae''
Cory As a given name, Cory is used by both males and females. It is a variation of the name Cora, meaning "(the) Maiden", which is a title of the goddess Persephone. The name also can have origins from the Gaelic word ''coire'', which means "in a cauldr ...
, 1916
The
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
partially separates ''T. c. sclateri'' as "common tody-flycatcher (''sclateri'') within the species; the other seven subspecies are the "common tody-flycatcher (''cinereum'' group)".Clements, J. F., P.C. Rasmussen, T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, A. Spencer, S. M. Billerman, B. L. Sullivan, M. Smith, and C. L. Wood. 2024. The eBird/Clements checklist of birds of the world: v2024. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved October 23, 2024


Description

The common tody-flycatcher is a tiny, big-headed bird, long, weighing , and with a long, flattened, straight bill. Adult males of the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
''T. c. cinereum'' have a glossy black forecrown and a slate-gray hindcrown and nape. Their lores and the area around their eyes are glossy black. Their upper back is slate-gray that becomes dark olive all the way to the uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. Their wings are black with yellow edges on the flight feathers and yellow edges and tips on the coverts; the last show as two
wing bar The following is a glossary of common English language terms used in the description of birds—warm-blooded vertebrates of the class Aves and the only living dinosaurs. Birds, who have and the ability to (except for the approximately 60 ext ...
s. Their tail is black with white tips on the outer feathers. Their entire underparts are bright yellow. They have an all-black
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
and a black
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
with a pinkish white base. Adult females have a grayer head than males and an entirely pinkish white mandible. Both sexes have a yellowish to white iris and bluish gray legs and feet. Juveniles have dark gray crown and cheeks, a buffy tinge to the wings' yellow parts, a dark iris, and paler yellow underparts than adults. The other subspecies of the common tody-flycatcher differ from the nominate and each other thus: * ''T. c. virididorsale'': brighter green upperparts than nominate * ''T. c. finitimum'': grayish (somewhat slaty) green upperparts * ''T. c. wetmorei'': brighter green upperparts than nominate * ''T. c. sclateri'': white throat and sometimes a dark iris * ''T. c. peruanum'': like nominate but with a dark iris * ''T. c. coloreum'': paler gray nape and more olive upperparts than nominate * ''T. c. cearae'': paler gray nape and more olive upperparts than nominate


Distribution and habitat

The common tody-flycatcher is found from Mexico to northern Argentina and southern Brazil, though its range does not include most of the
Amazon Basin The Amazon basin is the part of South America drained by the Amazon River and its tributary, tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries ...
. The subspecies are found thus: * ''T. c. virididorsale'': central
Veracruz Veracruz, formally Veracruz de Ignacio de la Llave, officially the Free and Sovereign State of Veracruz de Ignacio de la Llave, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entit ...
and adjacent northern
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
in southern Mexico * ''T. c. finitimum'': from southern Veracruz,
Chiapas Chiapas, officially the Free and Sovereign State of Chiapas, is one of the states that make up the Political divisions of Mexico, 32 federal entities of Mexico. It comprises Municipalities of Chiapas, 124 municipalities and its capital and large ...
,
Tabasco Tabasco, officially the Free and Sovereign State of Tabasco, is one of the Political divisions of Mexico, 32 Federal Entities of Mexico. It is divided into Municipalities of Tabasco, 17 municipalities and its capital city is Villahermosa. It i ...
, and the
Yucatán Peninsula The Yucatán Peninsula ( , ; ) is a large peninsula in southeast Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north and west of the peninsula from the C ...
in southern Mexico south on both the Caribbean and Pacific slopes to northwestern Costa Rica * ''T. c. wetmorei'': central and eastern Costa Rica and Panama;
Coiba Coiba () is the largest island in Central America, with an area of , off the Pacific coast of the Panamanian province of Veraguas. It is part of the Montijo District of that province. History Coiba separated from continental Panama between 1 ...
Island off Panama's Pacific coast * ''T. c. sclateri'': from Cauca and Nariño departments in southwestern Colombia south through western Ecuador to
Lambayeque Department Lambayeque () is a department and region in northwestern Peru known for its rich Moche and Chimú historical past. The region's name originates from the ancient pre-Inca civilization of the '' Lambayeque (''muchic: ''Ñampaxllæc)'' . It is ...
in far northwestern Peru * ''T. c. cinereum'': Colombia's three
Andean The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long and wide (widest between 18°S ...
ranges (except in the southwest) and from
Meta Department Meta () is a department of Colombia. It is close to the geographic center of the country, to the east of the Andean mountains. A large portion of the department, which is also crossed by the Meta River, is covered by a grassland plain known ...
north and east across northern Venezuela except its far northwest,
the Guianas The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British, Dutch, and French Guiana respectiv ...
, and northeastern Brazil from eastern
Roraima Roraima ( ) is one of the 26 states of Brazil. Located in the country's North Region, it is the northernmost and most geographically and logistically isolated state in Brazil. It is bordered by the state of Pará to the southeast, Amazonas t ...
to the Atlantic in
Amapá Amapá (; ) is one of the 26 federative units of Brazil, states of Brazil. It is in the North Region, Brazil, North Region of Brazil. It is Federative units of Brazil#List, the second-least populous state and the eighteenth-largest state by area ...
* ''T. c. peruanum'': eastern Ecuador and south in eastern Peru to Cuzco Department * ''T. c. coloreum'': northern Bolivia, northern Paraguay, far northeastern Argentina's
Misiones Province Misiones (, ''Missions'') is one of the Provinces of Argentina, 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamia, Argentina, Mesopotamia region. It is surrounded by Paraguay to the northwest, Brazil ...
, and southeastern Brazil roughly bounded by southern
Mato Grosso Mato Grosso ( – ) is one of the states of Brazil, the List of Brazilian states by area, third largest by area, located in the Central-West Region, Brazil, Central-West region. The state has 1.66% of the Brazilian population and is responsible ...
,
Espírito Santo Espírito Santo (; ) is a state in southeastern Brazil. Its capital is Vitória, and its largest city is Serra. With an extensive coastline, the state hosts some of the country's main ports, and its beaches are significant tourist attracti ...
, and Santa Catarina states * ''T. c. cearae'': eastern Brazil roughly bounded by
Pará Pará () is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian st ...
,
Alagoas Alagoas () is one of the 27 federative units of Brazil and is situated in the eastern part of the Northeast Region, Brazil, Northeast Region. It borders: Pernambuco (N and NW); Sergipe (S); Bahia (SW); and the Atlantic Ocean (E). Its capital is ...
, and northern
Bahia Bahia () is one of the 26 Federative units of Brazil, states of Brazil, located in the Northeast Region, Brazil, Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo (state), São Paulo, Mina ...
The common tody-flycatcher inhabits a wide variety of open and semi-open landscapes including
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
, forest edges, mangroves, riverine belts, open woodlands and groves, plantations and orchards,
restinga Restingas () are a distinct type of coastal tropical and subtropical moist broadleaf forest in eastern Brazil. They form on sandy, acidic, and nutrient-poor soils, and are characterized by medium-sized trees and shrubs adapted to the drier and n ...
, thickets in savanna, overgrown clearings and pastures, agricultural areas, and gardens. It shuns dense forest. In elevation it ranges from sea level to in northern Central America and in Costa Rica. It reaches in Colombia and in western Ecuador. It ranges between in eastern Ecuador and in Peru. In Venezuela it reaches north of the
Orinoco River The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers approximately 1 million km2, with 65% of it in Venezuela and 35% in Colombia. It is the List of rivers by discharge, f ...
and south of it. In Brazil it ranges from sea level to .


Behavior


Movement

The common tody-flycatcher is believed to be a year-round resident.


Feeding

The common tody-flycatcher feeds on a wide variety of
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s and also includes fruit in its diet. It typically forages in pairs or in small family groups, though sometimes singly, and rarely joins
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. It mostly forages in dense vegetation near the ground but sometimes ascends as high as the canopy where the forest is most open. It primarily takes prey from foliage by gleaning while perched and with short upward and outward sallies from a perch. It occasionally captures prey in mid-air. It forages very actively, hopping sideways along branches with its tail cocked and wagging. A study in Brazil showed that there it took almost all of its prey from live leaves by using strikes from a sally and hover-gleaning about equally.


Breeding

The common tody-flycatcher's breeding season varies geographically but overall is between December and October. Both sexes build the nest, a messy pouch with a side entrance under a "visor". It is made from grasses, other plant fibers, and moss bound with spider web and lined with fine grass, feathers, or seed down. Often fibers hang messily from the nest's bottom. It is typically suspended from a branch between about above the ground, sometimes near a wasp nest. Nests have also been recorded as high as and have been see hanging from utility wires. The usual clutch is two or three eggs which the female alone incubates. The incubation period is about 18 days and fledging occurs about 17 to 18 days after hatch. Both parents provision nestlings and fledglings.


Vocal and non-vocal sounds

The common tody-flycatcher's vocalizations include a "series of low, short, dry 'trrr-trrr' trills, a variety of high/very high, single, double, and triple 'tic' notes and nextr. high 'weet-widiwi' (last part as almost-trill)". The 'tic' notes can be "repeated up to 110 times per minute at dawn during henesting season". Prey capture is accompanied by an audible bill snap.


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the common tody-flycatcher as being of Least Concern. It has an extremely large range; its estimated population of at least five million mature individuals is believed to be stable. No immediate threats have been identified. It occurs in many protected areas both public and private and " rvives well in converted and secondary habitats; benefits from deforestation, spreading into clearings and plantations, where it is usually common". It is considered fairly common to common in northern Central America, common in most of Costa Rica though less so in the far northwest, "generally numerous" in Ecuador, and common in Colombia, Peru, Venezuela, and Brazil.


References


Further reading

*


External links

* {{Authority control
common tody-flycatcher The common tody-flycatcher (''Todirostrum cinereum'') is a small passerine bird in the family Tyrannidae, the tyrant flycatchers. It is found in Mexico, in every Central American country, and in every mainland South American country except Ch ...
Birds of Central America Birds of Brazil Birds of Bolivia Birds of Ecuador Birds of Colombia Birds of Venezuela Birds of Paraguay Birds of the Guiana Shield Birds of Peruvian Amazonia
common tody-flycatcher The common tody-flycatcher (''Todirostrum cinereum'') is a small passerine bird in the family Tyrannidae, the tyrant flycatchers. It is found in Mexico, in every Central American country, and in every mainland South American country except Ch ...
common tody-flycatcher The common tody-flycatcher (''Todirostrum cinereum'') is a small passerine bird in the family Tyrannidae, the tyrant flycatchers. It is found in Mexico, in every Central American country, and in every mainland South American country except Ch ...