''Crangon crangon'' is a species of
caridean shrimp found across the northeastern
Atlantic Ocean
The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
. Its range extends from the
White Sea
The White Sea (; Karelian language, Karelian and ; ) is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is surrounded by Karelia to the west, the Kola Peninsula to the north, and the Kanin Peninsula to the nort ...
in the north of
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
to the coast of
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, including the
Baltic Sea
The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by the countries of Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden, and the North European Plain, North and Central European Plain regions. It is the ...
, and appears also throughout the
Mediterranean
The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
and
Black Sea
The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
s.
Commercially important, it is fished mainly in the southern
North Sea
The North Sea lies between Great Britain, Denmark, Norway, Germany, the Netherlands, Belgium, and France. A sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian Se ...
.
Common name
In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s include brown shrimp,
common shrimp, bay shrimp, and sand shrimp, while translation of its French name ' (or its Dutch equivalent ') sometimes leads to the English version grey shrimp. Krangṓn (κραγγών) is an
Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
word for shrimp.
Description
Adults are typically long, although individuals up to have been recorded.
The animals have
cryptic colouration, being a sandy brown colour, which can be changed to match the environment.
They live in shallow water, which can also be slightly
brackish
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
, and feed nocturnally.
During the day, they remain buried in the sand to escape predatory birds and fish, with only their
antennae protruding.
''Crangon'' is classified in the family
Crangonidae
Crangonidae is a family of shrimp, of the superfamily Crangonoidea, including the commercially important species ''Crangon crangon''. Its type genus is '' Crangon''. Crangonid shrimps' first pair of pereiopods have partially chelate claws tha ...
, and shares the family's characteristic subchelate first
pereiopod
The anatomy of a decapod consists of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment – often called a somite – may possess one pair of appendages, although in various groups these m ...
s (where the movable finger closes onto a short projection, rather than a similarly sized fixed finger) and short
rostrum
Rostrum may refer to:
* Any kind of a platform for a speaker:
**dais
**pulpit
** podium
* Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects
* Rostrum (ship), a form of bow on naval ...
.
Distribution and ecology
''C. crangon'' has a wide
range
Range may refer to:
Geography
* Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra)
** Mountain range, a group of mountains bordered by lowlands
* Range, a term used to i ...
, extending across the northeastern
Atlantic Ocean
The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
from the
White Sea
The White Sea (; Karelian language, Karelian and ; ) is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is surrounded by Karelia to the west, the Kola Peninsula to the north, and the Kanin Peninsula to the nort ...
in the north of
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
to the coast of
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, including the
Baltic Sea
The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by the countries of Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden, and the North European Plain, North and Central European Plain regions. It is the ...
, as well as occurring throughout the
Mediterranean
The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
and
Black Sea
The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
s. Despite its wide range, however, little
gene flow
In population genetics, gene flow (also known as migration and allele flow) is the transfer of genetic variation, genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent ...
occurs across certain natural barriers, such as the
Strait of Gibraltar
The Strait of Gibraltar is a narrow strait that connects the Atlantic Ocean to the Mediterranean Sea and separates Europe from Africa.
The two continents are separated by 7.7 nautical miles (14.2 kilometers, 8.9 miles) at its narrowest point. Fe ...
or the
Bosphorus
The Bosporus or Bosphorus Strait ( ; , colloquially ) is a natural strait and an internationally significant waterway located in Istanbul, Turkey. The Bosporus connects the Black Sea to the Sea of Marmara and forms one of the continental bo ...
.
The populations in the western Mediterranean Sea are thought to be the oldest, with the species' spread across the north Atlantic thought to postdate the
Pleistocene
The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
.
Adults live epibenthically (on or near the sea-floor) especially in the shallow waters of estuaries or near the coast.
It is generally highly abundant, and has a significant effect on the ecosystems where it lives.
Lifecycle
Females reach
sexual maturity
Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
at a length around , while males are mature at .
The young hatch from their eggs into
plankton
Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
ic
larvae
A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect developmental biology, development such as insects, some arachnids, amphibians, or cnidarians typical ...
. These pass through five
moults before reaching the postlarval stage, when they settle to the sea-floor.
Fishery

Historically, the commercial fishery was accomplished by horse-drawn beam trawls on both sides of the Dover straits. In the sandy shallows of
Morecambe Bay
Morecambe Bay is an estuary in north-west England, just to the south of the Lake District National Park. It is the largest expanse of intertidal mudflats and sand in the United Kingdom, covering a total area of . In 1974, the second largest ga ...
(Lancashire, UK) horses have been replaced by tractors. Some small fishing vessels also use beam trawls for brown shrimp. A few artisanal fishermen use hand-pushed nets. In all UK shrimp fisheries, the catch is first 'riddled' to release the young of shrimps and fish. The shrimps are then traditionally boiled on board before landing.
Over of ''C. crangon'' were caught in 1999, with
Germany
Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
and the
Netherlands
, Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
taking over 80% of this total.
The UK lands an annual average of 1000 tonnes of brown shrimp, but the catch is highly variable between 500 and 1500 tonnes.
In the Lancashire fishery for brown shrimp it has been shown that landings in any year are related to the annual catch, average annual air temperature (inverse) and total rainfall in the previous year. That has enabled a good prediction of annual landings one year in advance. Moreover, for the port of Lytham, the abundance of shrimp (annual catch per unit effort) was found to be closely correlated with the mean annual
Zürich sunspot number for the period 1965-1975. Given that sunspot numbers are predictable, this provides another tool for the prediction of annual shrimp catch. Sunspot cycle No. 23 (1997–2008) is a good example of the correlation between UK annual brown shrimp catch and mean annual sunspot number.
Greenpeace
Greenpeace is an independent global campaigning network, founded in Canada in 1971 by a group of Environmental movement, environmental activists. Greenpeace states its goal is to "ensure the ability of the Earth to nurture life in all its biod ...
Germany classifies the brown shrimp as an "unsustainable" choice that should be avoided. Brown shrimp have been documented to contain
microplastics
Microplastics are "synthetic solid particles or polymeric matrices, with regular or irregular shape and with size ranging from 1 μm to 5 mm, of either primary or secondary manufacturing origin, which are insoluble in water." Microplastics a ...
.
As food

The consumption of brown shrimp is popular in Belgium, the Netherlands, northern Germany, and Denmark.
Shrimp in general are known as ''garnalen'' in
Dutch. It is the basis of the dish ''
tomate-crevettes'', where the shrimp are mixed with
mayonnaise
Mayonnaise (), colloquially referred to as "mayo" (), is a thick, creamy sauce with a rich and tangy taste that is commonly used on sandwiches, hamburgers, Salad#Bound salads, bound salads, and French fries. It also forms the base for various o ...
and fresh
parsley
Parsley, or garden parsley (''Petroselinum crispum''), is a species of flowering plant in the family Apiaceae that is native to Greece, Morocco and the former Yugoslavia. It has been introduced and naturalisation (biology), naturalized in Eur ...
, and served in a hollowed-out uncooked
tomato
The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
. The shrimp
croquette
A croquette (; ) is a deep-fried roll originating in French cuisine, consisting of a thick binder combined with a filling, which is then breaded. It is served as a side dish, a snack, or fast food worldwide.
The binder is typically a thick b� ...
is another Belgian speciality; the shrimp are in the interior of the battered croquette along with
béchamel sauce. Freshly cooked, unpeeled brown shrimp are often served as a snack accompanying beer, typically a sour ale or
Flemish red such as
Rodenbach.
In
Lancashire
Lancashire ( , ; abbreviated ''Lancs'') is a ceremonial county in North West England. It is bordered by Cumbria to the north, North Yorkshire and West Yorkshire to the east, Greater Manchester and Merseyside to the south, and the Irish Sea to ...
, England, the peeled brown shrimps are mixed with butter and spices (including nutmeg or mace) to make
potted shrimps, a dish traditionally eaten with bread.
References
External links
*
*
{{Portal bar, Crustaceans, Marine life
Caridea
Commercial crustaceans
Crustaceans described in 1758
Crustaceans of Europe
Caridea of the Atlantic Ocean
Edible crustaceans
Extant Pleistocene first appearances
Quaternary animals of Europe
Taxa named by Carl Linnaeus
Crangonidae