Common Scale-backed Antbird
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The common scale-backed antbird (''Willisornis poecilinotus'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird in subfamily Thamnophilinae of family
Thamnophilidae The antbirds are a large passerine bird family, Thamnophilidae, found across subtropical and tropical Central and South America, from Mexico to Argentina. There are more than 230 species, known variously as antshrikes, antwrens, antvireos, fire ...
, the "typical antbirds". It is found in
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
,
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
,
French Guiana French Guiana, or Guyane in French, is an Overseas departments and regions of France, overseas department and region of France located on the northern coast of South America in the Guianas and the West Indies. Bordered by Suriname to the west ...
,
Guyana Guyana, officially the Co-operative Republic of Guyana, is a country on the northern coast of South America, part of the historic British West Indies. entry "Guyana" Georgetown, Guyana, Georgetown is the capital of Guyana and is also the co ...
,
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
,
Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...
, and
Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...
.


Taxonomy and systematics

The common scale-backed antbird has a complicated taxonomic history. It was described and illustrated by the German ornithologist
Jean Cabanis Jean Louis Cabanis (8 March 1816 – 20 February 1906) was a German ornithologist. He worked at the bird collections of the Natural History Museum in Berlin becoming its first curator of birds in 1850. He founded the ''Journal für Ornithologie ...
in 1847 and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Hypocnemis poecilinotus''. The
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
is from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''poikilonōtos'' "with variegated back" (from ''poikilos'' "spotted" and ''nōton'' "back"). It was subsequently included in the genus '' Hylophylax'', but was found to not be closely related to the other species in the genus and then was briefly placed in genus ''Dichropogon''. This name is preoccupied by a genus of asilid flies (''Dichropogon'' Bezzi, 1910) so the current genus ''Willisornis'' was created for it. The common scale-backed antbird has these five subspecies: * ''W. p. poecilinotus'' (
Cabanis Cabanis is a surname. Notable people with the surname include: * Bernard Cabanis (born 1950), French ice hockey player * George Cabanis (1815–1892), American politician * James Cabanis (1838–1920), American politician, son of George Cabanis *Jea ...
, 1847)
* ''W. p. duidae'' ( Chapman, 1923) * ''W. p. lepidonota'' ( Sclater, PL & Salvin, 1880) * ''W. p. griseiventris'' ( Pelzeln, 1868) * ''W. p. gutturalis'' (
Todd Todd or Todds may refer to: Places Australia * Todd River, an ephemeral river United States * Todd Valley, California, also known as Todd, an unincorporated community * Todd, Missouri, a ghost town * Todd, North Carolina, an unincorporated c ...
, 1927)
The
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
s of the subspecies differ greatly, leading to speculation that some of them should be recognized as full species. The South American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
is seeking a formal proposal to consider the issue.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, G. Del-Rio, A. Jaramillo, D. F. Lane, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 4 March 2024. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved March 5, 2024 What are now the two subspecies of the
Xingu scale-backed antbird The Xingu scale-backed antbird (''Willisornis vidua'') is a species of bird in subfamily Thamnophilinae of family Thamnophilidae, the "typical antbirds". It is endemic to Brazil. Taxonomy and systematics What is now the Xingu scale-backed an ...
(''W. vidua'') were previously included in ''W. poecilinotus''.


Description

The common scale-backed antbird is long and weighs . Adult males of the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
''W. p. poecilinotus'' are mostly gray; their upperparts are darker than their underparts. They have a white patch between the
scapulars The scapula (: scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side ...
. Their lower back has white-edged black patches that give it its English name. Their wings and tail are black with white edges and tips on many feathers. Adult females have a reddish yellow-brown crown and face. Their upperparts are mostly olive-brown with buff-edged brownish black patches on the lower back. They also have a white patch between the scapulars. Their flight feathers are blackish brown with reddish brown edges and their tail is dark olive-brown with white spots and a blackish band near the end. Their throat is pale grayish white. Their underparts are gray with a reddish yellow-brown tinge on the flanks.Zimmer, K., M.L. Isler, and G. M. Kirwan (2020). Common Scale-backed Antbird (''Willisornis poecilinotus''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.scbant3.01 retrieved June 13, 2024 Females of subspecies ''W. p. duidae'' have a black lower back and rump with wide white feather edges, a white-spotted black tail, reddish yellow-brown underparts with a cinnamon-rufous center to the belly, and olive-brown tinged flanks. Females of ''W. p. lepidonota'' are similar to those of ''duidae'' but paler and with a browner crown and a light buff center to the belly. Females of ''W. p. griseiventris'' have a rufous-buff face, a plain lower back and rump, a pale gray throat, and gray underparts. Males of ''W. p. gutturalis'' have a black throat; females resemble female ''lepidonota'' but with a pale olive center to the belly.Schulenberg, T.S., D.F. Stotz, D.F. Lane, J.P. O’Neill, and T.A. Parker III. 2010. ''Birds of Peru''. Revised and updated edition. Princeton University Press, Princeton, New Jersey. Plate 175


Distribution and habitat

The subspecies of the common scale-backed antbird are found thus: * ''W. p. poecilinotus'': central and southeastern Venezuela, the
Guiana The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British, Dutch, and French Guiana respectiv ...
s, and northeastern Brazil from
Roraima Roraima ( ) is one of the 26 states of Brazil. Located in the country's North Region, it is the northernmost and most geographically and logistically isolated state in Brazil. It is bordered by the state of Pará to the southeast, Amazonas t ...
to
Amapá Amapá (; ) is one of the 26 federative units of Brazil, states of Brazil. It is in the North Region, Brazil, North Region of Brazil. It is Federative units of Brazil#List, the second-least populous state and the eighteenth-largest state by area ...
* ''W. p. duidae'': east-central Colombia, southwestern Venezuela, and northwestern Brazil along the Rio Negro * ''W. p. lepidonota'': southeastern Colombia, eastern Ecuador, and northeastern and east-central Peru * ''W. p. griseiventris'': south-central and southeastern Peru, northern Bolivia, and southwestern Amazonian Brazil between the rios Juruá and
Madeira Madeira ( ; ), officially the Autonomous Region of Madeira (), is an autonomous Regions of Portugal, autonomous region of Portugal. It is an archipelago situated in the North Atlantic Ocean, in the region of Macaronesia, just under north of ...
south into western
Mato Grosso Mato Grosso ( – ) is one of the states of Brazil, the List of Brazilian states by area, third largest by area, located in the Central-West Region, Brazil, Central-West region. The state has 1.66% of the Brazilian population and is responsible ...
* ''W. p. gutturalis'': northeastern Peru and western Amazonian Brazil to the Rio Juruá The common scale-backed antbird primarily inhabits the understorey of humid '' terra firme''
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, live oak, acacias, magnolia, and banksia in more temperate zo ...
. It also occurs in '' várzea'' and ''
igapó (, from Tupi language, Old Tupi: "root forest") is a word used in Brazil for Blackwater river, blackwater-flooded forests in the Amazon biome. These forests and similar swamp forests are seasonally inundated with freshwater. They typically occur ...
'' forest. In elevation it reaches in Brazil, in Venezuela, and in Colombia. In Ecuador it mostly occurs below but does reach . In Peru it occurs locally to but is mostly below .


Behavior


Movement

The common scale-backed antbird is believed to be a year-round resident throughout its range.


Feeding

The common scale-backed antbird feeds mostly on a wide variety of
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s; it has also been noted eating small lizards. It typically forages individually, in pairs, and in family groups, usually within about of the ground though sometimes as high as and rarely to . In some parts of its range it joins
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. It follows swarms of
army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limited ...
s that pass through its territory to capture prey fleeing the ants, but it just as often forages away from ant swarms. It captures prey mostly by short sallies from a perch to the ground but also to foliage, branches, and vines. It sometimes stays on the ground probing and flicking leaf litter. At ant swarms it defers to other species.


Breeding

The breeding season of the common scale-backed antbird varies greatly across its range, for instance spanning November to March in French Guiana and probably December to July in Ecuador. It possibly breeds in any month in Brazil. It has been documented building several types of nests, including an open cup of dry leaves and other fibers in an understorey palm, a cavity lined with dead leaves, an unlined hollow at the top of a stump, and a cup of dried grass on the ground. The usual clutch is two eggs. The female alone is believed to incubate at night but both parents do so during the day. Both parents also brood and provision nestlings. The incubation period and time to fledging are not known.


Vocalization

The common scale-backed antbird's songs are generally " a series...of long upslurred notes...with little space between them, each note rising in pitch and gaining in intensity, except final note or notes that decrease in intensity". The number of notes and some other features differ among individuals and also among the subspecies. In Venezuela it has been written as "''pureeeee, pureeeee, pureeeee...''", in Ecuador as "teeuw, tuweeé? tuweeé? tuweeé? tuweeé?", and in Peru as "''hew hui hui hui? hui? hui? hui? hui? hui?''". Its calls include a "sharp 'psit'...a usually descending 'chirr', more abrupt chitter, short whistle quickly repeated 3–4 times, ndalso a longer whistle that falls and then rises in pitch, sounding like 'cherri' ".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the common scale-backed antbird as being of Least Concern. It has a very large range. Its population size is not known and is believed to be stable. No immediate threats have been identified. It is considered fairly common throughout its range, which includes many large protected areas. "Regions occupied by this species also encompass extensive intact habitat which, although not formally protected, seems to be at little near-term risk."


References


External links


Scale-backed antbird photo gallery
VIREO
Photo
chandra.as.utexas.edu {{Taxonbar, from=Q1063238 Willisornis Birds of the Amazon rainforest Birds of the Guiana Shield Birds described in 1847 Taxonomy articles created by Polbot