Common Aspen
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''Populus tremula'' (commonly called aspen, common aspen, Eurasian aspen, European aspen, or quaking aspen) is a species of poplar native to cool
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
regions of the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
.


Description

It is a substantial
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
growing to tall by broad, with a trunk attaining over in diameter. The
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
is pale greenish-grey and smooth on young trees with dark grey diamond-shaped
lenticel A lenticel is a porous tissue consisting of cells with large intercellular spaces in the periderm of the secondarily thickened organs and the Bark (botany), bark of woody stems and roots of gymnosperms and dicotyledonous flowering plants. It func ...
s, becoming dark grey and fissured on older trees. The adult
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
, produced on branches of mature trees, are nearly round, slightly wider than long, diameter, with a coarsely toothed margin and a laterally flattened
petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...
long. The flat petiole allows them to tremble in even slight breezes, and is the source of its scientific name, as well as one of its vernacular names "langues de femmes" attributed to Gerard's 17th-century ''Herball''. The leaves on seedlings and fast-growing stems of suckers (root sprouts) are of a different shape, heart-shaped to nearly triangular. They are also often much larger, up to long; their petiole is also less flattened. The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are wind-pollinated
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind- pollinated ( anemophilous) but sometimes insect-pollinated (as in '' Salix''). It contains many, usually unisexual flowers, arra ...
s produced in early spring before the new leaves appear; they are
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
, with male and female catkins on different trees. The male catkins are patterned green and brown, long when shedding pollen; the female catkins are green, long at pollination, maturing in early summer to bear 10–20 (50–80) capsules each containing numerous tiny
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s embedded in downy fluff. The fluff assists wind dispersal of the seeds when the capsules split open at maturity.Trees for Life Species Profile
Aspen
Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins . It can be distinguished from the closely related North American ''
Populus tremuloides ''Populus tremuloides'' is a deciduous tree native to cooler areas of North America, one of several species referred to by the common name aspen. It is commonly called quaking aspen,
,'' which is nearly identical, by the leaves being more coarsely toothed. Like other aspens, it spreads extensively by suckers (root sprouts), which may be produced up to 40 m from the parent tree, forming extensive
clonal colonies A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in ...
. This often makes the job of clearing unwanted trees from an area especially difficult, as new suckers will continue to sprout from the extensive root system for up to several years after all surface growth has been eliminated.


Distribution and habitat

The species is native to Europe and Asia, from Iceland and the British IslesJames Kilkelly east to
Kamchatka The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...
, north to inside the
Arctic Circle The Arctic Circle is one of the two polar circles, and the northernmost of the five major circle of latitude, circles of latitude as shown on maps of Earth at about 66° 34' N. Its southern counterpart is the Antarctic Circle. The Arctic Circl ...
in Scandinavia and northern Russia, and south to central Spain, Turkey, the
Tian Shan The Tian Shan, also known as the Tengri Tagh or Tengir-Too, meaning the "Mountains of God/Heaven", is a large system of mountain ranges in Central Asia. The highest peak is Jengish Chokusu at high and located in Kyrgyzstan. Its lowest point is ...
, North Korea, and northern Japan. It also occurs at one site in
northwest Africa The Maghreb (; ), also known as the Arab Maghreb () and Northwest Africa, is the western part of the Arab world. The region comprises western and central North Africa, including Algeria, Libya, Mauritania, Morocco, and Tunisia. The Maghreb al ...
in
Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
. In the south of its range, it occurs at high altitudes in mountains.Den Virtuella Floran
''Populus tremula'' (in Swedish; with maps)
/ref>


Ecology

Eurasian aspen is a water and light demanding species that is able to vigorously colonize an open area after fire,
clear cutting Clearcutting, clearfelling or clearcut logging is a forestry/logging practice in which most or all trees in an area are uniformly cut down. Along with Shelterwood cutting, shelterwood and Seed tree, seed tree harvests, it is used by foresters t ...
or other kinds of damage. After an individual has been damaged or destroyed, root suckers are produced abundantly on the shallow
lateral roots Lateral roots, emerging from the pericycle (meristematic tissue), extend horizontally from the primary root (radicle) and over time makeup the iconic branching pattern of root systems. They contribute to anchoring the plant securely into the soil ...
. Fast growth continues until the age of about 20 years, when crown competition increases. After that, growth speed decreases and culminates at about 30 years of age. Aspen can reach an age of 200 years. It is a very hardy species and tolerates long, cold winters and short summers. Aspen is resistant to browsing pressure by
fallow deer Fallow deer is the common name for species of deer in the genus ''Dama'' of subfamily Cervinae. There are two living species, the European fallow deer (''Dama dama''), native to Europe and Anatolia, and the Persian fallow deer (''Dama mesopotamic ...
owing to its unpleasant taste. This species is important for the
hornet moth The hornet moth or hornet clearwing (''Sesia apiformis'') is a large moth native to Europe and the Middle East and has been introduced to North America. Its protective coloration is an example of Batesian mimicry, Batesian mimicry, as its simil ...
, which uses it as a host during the larval stage.


Fossil record

Fossils of ''Populus tremula'' have been described from the
fossil flora of Kızılcahamam district This fossil flora in Turkey stems from at least six Pliocene deposits in Güvem and Beşkonak villages, north of Ankara, north of Kızılcahamam and south of the Black Sea coast. They are to above sea level, extending north-south and eas ...
in
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
which is of early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58fastigiate This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary ...
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
'Erecta', with bright yellow autumn colouring, has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
. The cultivar is colloquially known as "Swedish columnar" in Canada and the United States. The
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
with ''
Populus alba ''Populus alba'', commonly called silver poplar,Webb, C. J.; Sykes, W. R.; Garnock-Jones, P. J. 1988: Flora of New Zealand. Vol. IV. Naturalised Pteridophytes, Gymnosperms, Dicotyledons. 4. Christchurch, New Zealand, Botany Division, D.S.I.R. si ...
'' (white poplar), known as grey poplar, ''Populus'' × ''canescens'', is widely found in Europe and central Asia. Hybrids with several other aspens have also been bred at forestry research institutes in order to find trees with greater timber production and disease resistance (e.g. ''P. tremula'' × ''P. tremuloides'', bred in DenmarkJensen, N. (1994). Guide til Arboretet i Hørsholm (in Danish).).


Use

The wood of aspen is light and soft with very little shrinkage. It is used for
lumber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
and
matches A match is a tool for starting a fire. Typically, matches are made of small wooden sticks or stiff paper. One end is coated with a material that can be ignited by friction generated by striking the match against a suitable surface. Wooden matc ...
but is also valued in the
pulp and paper industry The pulp and paper industry comprises companies that use wood, specifically pulpwood, as raw material and produce pulp, paper, paperboard, and other cellulose-based products. Manufacturing process In the manufacturing process, pulp is intr ...
, being particularly useful for
writing paper Printing and writing papers are paper grades used for newspapers, magazines, catalogs, books, notebooks, commercial printing, business forms, stationeries, copying and digital printing. About 1/3 of the total pulp and paper marked (in 2000) is p ...
. In addition, it is used for
plywood Plywood is a composite material manufactured from thin layers, or "plies", of wood veneer that have been stacked and glued together. It is an engineered wood from the family of manufactured boards, which include plywood, medium-density fibreboa ...
and different types of flake and particle boards. Given its hardiness and capacity for rapid growth and regeneration, it plays an important role in the production of wood for
renewable energy Renewable energy (also called green energy) is energy made from renewable resource, renewable natural resources that are replenished on a human lifetime, human timescale. The most widely used renewable energy types are solar energy, wind pow ...
. Ecologically, the species is important as many insect and fungi species benefit from it. The tree further provides
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
for several mammals and birds that require young forests.


References


External links


Video, commentary and annotation on why Aspen leaves tremble.

''Populus tremula''
- distribution map, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that promotes the conservation and sustainable use of forest genetic resources in Europe as an integral part of sustainable forest management. It was established ...
(EUFORGEN) *
Populus tremula
' ''-'' Aspen; info, images and video a
Woodland Trust
* {{Authority control tremula Flora of temperate Asia Flora of Europe Flora of Algeria Trees of Europe Trees of continental subarctic climate Trees of humid continental climate Trees of subpolar oceanic climate Plants described in 1753 Taxa named by Carl Linnaeus