Combretum Glutinosum
   HOME

TheInfoList



OR:

''Combretum glutinosum'' is a shrub species of the genus ''
Combretum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar ...
'', found in the Sahel belt in parts of
Senegal Senegal, officially the Republic of Senegal, is the westernmost country in West Africa, situated on the Atlantic Ocean coastline. It borders Mauritania to Mauritania–Senegal border, the north, Mali to Mali–Senegal border, the east, Guinea t ...
,
Burkina Faso Burkina Faso is a landlocked country in West Africa, bordered by Mali to the northwest, Niger to the northeast, Benin to the southeast, Togo and Ghana to the south, and Ivory Coast to the southwest. It covers an area of 274,223 km2 (105,87 ...
,
Ghana Ghana, officially the Republic of Ghana, is a country in West Africa. It is situated along the Gulf of Guinea and the Atlantic Ocean to the south, and shares borders with Côte d’Ivoire to the west, Burkina Faso to the north, and Togo to t ...
,
Mali Mali, officially the Republic of Mali, is a landlocked country in West Africa. It is the List of African countries by area, eighth-largest country in Africa, with an area of over . The country is bordered to the north by Algeria, to the east b ...
,
the Gambia The Gambia, officially the Republic of The Gambia, is a country in West Africa. Geographically, The Gambia is the List of African countries by area, smallest country in continental Africa; it is surrounded by Senegal on all sides except for ...
,
Niger Niger, officially the Republic of the Niger, is a landlocked country in West Africa. It is a unitary state Geography of Niger#Political geography, bordered by Libya to the Libya–Niger border, north-east, Chad to the Chad–Niger border, east ...
,
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
and
Cameroon Cameroon, officially the Republic of Cameroon, is a country in Central Africa. It shares boundaries with Nigeria to the west and north, Chad to the northeast, the Central African Republic to the east, and Equatorial Guinea, Gabon, and the R ...
, across to parts of
Sudan Sudan, officially the Republic of the Sudan, is a country in Northeast Africa. It borders the Central African Republic to the southwest, Chad to the west, Libya to the northwest, Egypt to the north, the Red Sea to the east, Eritrea and Ethiopi ...
. It is known as in
Pulaar Pulaar (in Latin script, Latin: , in Ajami script, Ajami: ), often referred to as Pulaar du Nord, is dialect of the Fula language spoken primarily as a first language by the Fula people, Fula and Toucouleur peoples in the Senegal River valley ar ...
, ''Kantakara'' in
Hausa Hausa may refer to: * Hausa people, an ethnic group of West Africa * Hausa language, spoken in West Africa * Hausa Kingdoms, a historical collection of Hausa city-states * Hausa (horse) or Dongola horse, an African breed of riding horse See also ...
, ''rat'' in
Wolof Wolof or Wollof may refer to: * Wolof people, an ethnic group found in Senegal, Gambia, and Mauritania * Wolof language, a language spoken in Senegal, Gambia, and Mauritania * The Wolof or Jolof Empire, a medieval West African successor of the Mal ...
and ''jambakatan kè'' in
Maninka Maninka (also known as Malinke), or more precisely Eastern Maninka, is the name of several closely related languages and dialects of the southeastern Manding subgroup of the Mande language family (itself, possibly linked to the Niger–Congo ...
. Its synonyms are ''Combretum cordofanum'' Engl. & Diels, ''C. passargei'' Engl. & Diels, ''C. leonense'' Engl. & Diels.


Habitat

''Combretum glutinosum'' tends to grow in
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
type forests, in several soil types but is best suited to sandy and free draining soils. It is drought resistant and grows in areas of annual rainfall of . Its growth is fast and profuse.


Plant growth

As a bushy shrub the plant grows up to with an open crown with low branches that droop down, and is deciduous. The trunk is usually twisted and low branched, with grey-black rough bark. The thick leathery green leaves have a gummy feel to them and are glutinous when young. The plant flowers during the dry season after the bush fires which facilitate
leaf flushing Leaf flushing or leaf out is the production of a flush of new leaves typically produced simultaneously on all branches of a bare plant or tree. Young leaves often have less chlorophyll and the leaf flush may be white or red, the latter due to pre ...
that supports flowering. In Burkina Faso and Mali, flowers tend to appear between December and March, but this varies from area to area and flowers may appear as late as July. The fruit is a four-winged elliptic samara, and has a sticky feel, reddish and turns yellowish towards the end of the season. The fruit typically measures long and across. It bears fruits generally in January and fruiting lasts until November. Seeds are collected by shaking the branches of the tree. The seeds are oil bearing and one kilogram contains about 20,000 seeds, and it has an oil potential yield of c. 24%. The plant
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
which infects ''Combretum glutinosum'' is ''
Loranthaceae Loranthaceae, commonly known as the showy mistletoes, is a family of flowering plants. It consists of about 75 genera and 1,000 species of woody plants, many of them hemiparasites. The three terrestrial species are '' Nuytsia floribunda'' (the ...
''. Seeds stored for 18 months in temperature maintained at have a germination rate of 84% when planted. 95% germination has also been reported by "following drying to moisture contents in equilibrium with 15% relative humidity and freezing for 1 month at (-) 20 °C." The optimum temperature for germination is reported to be between and .


Chemical properties

Chemical analysis of the leaves of the plant has identified
Gallic acid Gallic acid (also known as 3,4,5-trihydroxybenzoic acid) is a trihydroxybenzoic acid with the formula C6 H2( OH)3CO2H. It is classified as a phenolic acid. It is found in gallnuts, sumac, witch hazel, tea leaves, oak bark, and other plant ...
,
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backward because it can be o ...
,
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
s and 4 tannins; the tannins reported are 2,3-(S)-hexahydroxydiphenoyl-D-glucose,
punicalin Punicalin is an ellagitannin. It can be found in '' Punica granatum'' (pomegranate) or in the leaves of '' Terminalia catappa'', a plant used to treat dermatitis and hepatitis. It is also reported in '' Combretum glutinosum'', all three species ...
,
punicalagin Punicalagin (''Pyuni-cala-jen'') is an ellagitannin, a type of phenolic compound. It is found as alpha and beta isomers in pomegranates (''Punica granatum''), '' Terminalia catappa'', '' Terminalia myriocarpa'', and in '' Combretum molle'', the ...
and combreglutinin. The black colour of the bogolan fabric is attributed to the chemical reaction of the tannins with the soluble iron compounds present in the fermented mud.


Uses

The bark, leaves and roots are used by the people inhabiting
Sahel The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
to produce yellow dye. The yellowish wood is "hard and extremely durable," and is used for building frames and tools, as well as fuel. The brownish or brownish yellow dyes are used in the textile industry and also in leather tanning and to dye mats. The plant is foraged by cattle,
giraffe The giraffe is a large Fauna of Africa, African even-toed ungulate, hoofed mammal belonging to the genus ''Giraffa.'' It is the Largest mammals#Even-toed Ungulates (Artiodactyla), tallest living terrestrial animal and the largest ruminant on ...
s and other animals, and its young leaves are sometimes eaten as a vegetable by humans, sometimes with
taro Taro (; ''Colocasia esculenta'') is a root vegetable. It is the most widely cultivated species of several plants in the family Araceae that are used as vegetables for their corms, leaves, stems and Petiole (botany), petioles. Taro corms are a ...
. The plant is in growing demand for the manufacture of bogolan textiles both in the local and international fashion world. ;Cultural A special textile known as the "mud cloth", "
bògòlanfini Bògòlanfini or bogolan (; "mud cloth"; sometimes called mud-dyed cloth or mud-painted cloth in English language, English) is a handmade Malian cotton fabric traditionally dyed with fermented mud. It has an important place in traditional Malian ...
" or " bogolan" is manufactured locally. It contains plant tannins mixed with iron
salts In chemistry, a salt or ionic compound is a chemical compound consisting of an assembly of positively charged ions ( cations) and negatively charged ions (anions), which results in a compound with no net electric charge (electrically neutral). ...
present in the fermented mud. Bogolan is a specialized art form established by Mande women, in particularly those from Bamanan, Bobo, Dogon, Malinke, Minianka, and Senoufo groups. ;Medicinal The plants bark, leaves and roots are extracted for traditional medicinal uses for treating various ailments from
influenza Influenza, commonly known as the flu, is an infectious disease caused by influenza viruses. Symptoms range from mild to severe and often include fever, runny nose, sore throat, muscle pain, headache, coughing, and fatigue. These sympto ...
, and
rheumatism Rheumatism or rheumatic disorders are conditions causing chronic, often intermittent pain affecting the joints or connective tissue. Rheumatism does not designate any specific disorder, but covers at least 200 different conditions, including a ...
, to sexual issues such as
impotence Erectile dysfunction (ED), also referred to as impotence, is a form of sexual dysfunction in males characterized by the persistent or recurring inability to achieve or maintain a Human penis, penile erection with sufficient rigidity and durat ...
and
syphilis Syphilis () is a sexually transmitted infection caused by the bacterium ''Treponema pallidum'' subspecies ''pallidum''. The signs and symptoms depend on the stage it presents: primary, secondary, latent syphilis, latent or tertiary. The prim ...
. It is commonly brewed as a tea in tropical West Africa to relieve stomach issues, and to treat malaria in a decoction with a number of other leaves obtained in the bush. In the Senegambia region, it is one of a number of trees whose twigs are used as "chewing sticks," used in lieu of toothbrushes to clean teeth, remove food particles after eating, and to chew for pleasure. It is also used to fill cavities of carious teeth, to dress wounds, for fumigation and as incense. A
decoction Decoction is a method of extraction by boiling herbal or plant material (which may include stems, roots, bark and rhizomes) to dissolve the chemicals of the material. It is the most common preparation method in various herbal medicine systems. D ...
of the leaves is used for baths to relieve fatigue.


References


External links


Further reading
{{Taxonbar, from=Q1113729 glutinosum Trees of Africa