The clouded salamander (''Aneides ferreus'') is a species of

salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...

in the family

Plethodontidae Plethodontidae, or lungless salamanders, are a family of salamanders. With over 500 species, lungless salamanders are by far the largest family of salamanders in terms of their diversity. Most species are native to the Western Hemisphere, from B ...

. It is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...

. Its natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

is temperate

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

s and it is probable that many nest in trees. It is threatened by

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

Description

The clouded salamander can grow to about total length. It is long and slim with relatively long legs, square ends to its toes and a prehensile tail. It has two naso-labial grooves joining its nostrils to its mouth and sixteen costal grooves down its flank. The upper side is pale grey variously blotched with gold, olive green or dull red. The juvenile has a brass-colored streak down the center of its back.

Distribution and habitat

The clouded salamander is found in the Cascadian bioregion of North America at heights up to . Its range extends from the

Columbia River The Columbia River (Upper Chinook language, Upper Chinook: ' or '; Sahaptin language, Sahaptin: ''Nch’i-Wàna'' or ''Nchi wana''; Sinixt dialect'' '') is the largest river in the Pacific Northwest region of North America. The river headwater ...

southwards through the

Cascade Mountains The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as many of those in the ...

, along the Oregon coast to the northern tip of California in

Del Norte County Del Norte County (Spanish for "Of The North") is a county located at the far northwest corner of the U.S. state of California, along the Pacific Ocean adjacent to the Oregon border. Its population was 27,743 as of the 2020 census, down from 28 ...

. It lives in forested areas favoring moist areas with few trees rather than dry dense stands. It hides under logs and rocks, under loose bark, in rotting logs and in crevices in rock and scree. It particularly favors rotting

Douglas-fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Or ...

logs. It is often common in newly cleared areas with tree stumps and debris.

Biology

The clouded salamander feeds on small invertebrates, mainly

woodlice Woodlice are terrestrial isopods in the suborder Oniscidea. Their name is derived from being often found in old wood, and from louse, a parasitic insect, although woodlice are neither parasitic nor insects. Woodlice evolved from marine isopods ...

ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...

s and

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

s, but also include

flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...

termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...

mite Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as eac ...

centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...

millipede Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...

spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...

s and

pseudoscorpion Pseudoscorpions, also known as false scorpions or book scorpions, are small, scorpion-like arachnids belonging to the order Pseudoscorpiones, also known as Pseudoscorpionida or Chelonethida. Pseudoscorpions are generally beneficial to humans bec ...

s. Breeding takes place in June and July with females laying clutches of nine to seventeen eggs, usually in chambers hollowed out in rotting logs. The larvae develop inside the eggs, emerging as juveniles in about two months, thus bypassing the free-living larval stage. One or both parents may provide care for the eggs before they hatch.

Status

The clouded salamander is listed as least concern in the

IUCN Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological spe ...

. This salamander's favored microhabitat of tree stumps and coarse woody decaying debris are being replaced by changes in forestry practices. However, plans to conserve the

spotted owl The spotted owl (''Strix occidentalis'') is a species of true owl. It is a resident species of old-growth forests in western North America, where it nests in tree hollows, old bird of prey nests, or rock crevices. Nests can be between high a ...

(''Strix occidentalis'') and the

marbled murrelet The marbled murrelet (''Brachyramphus marmoratus'') is a small seabird from the North Pacific. It is a member of the family Alcidae, which includes auklets, guillemots, murres and puffins. It nests in old-growth forests or on the ground at hig ...

(''Brachyramphus marmoratus'') may help to reduce the decline in numbers of clouded salamanders.

References

{{Authority control Aneides Amphibians described in 1869 Taxa named by Edward Drinker Cope Taxonomy articles created by Polbot