The clouded border (''Lomaspilis marginata'') is a

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

of the family

Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyleti ...

. The species was first described by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in his 1758 10th edition of ''Systema Naturae''. It is distributed across most of Europe to the

Urals The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.

, western and central

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

Transbaikalia Transbaikal, Trans-Baikal, Transbaikalia ( rus, Забайка́лье, r=Zabaykal'ye, p=zəbɐjˈkalʲjɪ), or Dauria (, ''Dauriya'') is a mountainous region to the east of or "beyond" (trans-) Lake Baikal at the south side of the eastern Si ...

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...

Tian-Shan The Tian Shan, also known as the Tengri Tagh or Tengir-Too, meaning the "Mountains of God/Heaven", is a large system of mountain ranges in Central Asia. The highest peak is Jengish Chokusu at high and located in Kyrgyzstan. Its lowest point is ...

, northern

Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...

and parts of the

Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...

Description

This is a very distinctive species with white wings marked with black blotches around the margins. The amount of black varies, with the males usually (though not always) having more extensive black areas than the females. Occasionally almost entirely white or black individuals are seen, although this is rare. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

is 24–28 mm. ''Lomaspilis marginata'' is extremely variable. Linnaeus's form has complete black border to both wings, also on the forewing additional spots or patches at base and middle of costa. Aberrations: * ab. ''nigrofasciata'' Schoyen: the median band, at least on the forewing, is complete, the marginal area is normal. * ab. ''mediofasciata'' Hofner: has a complete median band but no marginal band. * ab. ''dumeei'' de Joannis: lacks also the basal costal patch. * ab. ''suffusa''

rout A rout is a Panic, panicked, disorderly and Military discipline, undisciplined withdrawal (military), retreat of troops from a battlefield, following a collapse in a given unit's discipline, command authority, unit cohesion and combat morale ...

has the ground colour suffused with smoky yellow grey, the dark markings weak and blurred. * ab. ''nigrounicolorata'' Haverkampf: is black throughout. * ab. ''pollutaria'' Hbn.: has a narrow band of the ground colour at the distal margin, reducing and dividing into 2 parts the black marginal band. * ab. ''subdeleta'' Ckll: forewmg with marginal band almost or altogether obsolete, narrow patches at base and middle of costa remaining, hindwing unmarked. * ab. ''conflua'' Strand.: Costal patches confluent, also confluent with the marginal band. Median area normal. * ab: ''opis'' Btlr: has on each wing three large median spots, longer than broad. It forms ''opis'' a local race in S. E. Siberia and Japan." Wikisource:The Moths of the British Isles Second Series/Chapter 10#263 It flies at night in June and July and is attracted to light. The egg is yellow green, with hexagonal reticulation. The

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

, pale green with darker dorsal lines and a purplish anal spot, usually feeds on

aspen Aspen is a common name for certain tree species in the Populus sect. Populus, of the ''Populus'' (poplar) genus. Species These species are called aspens: * ''Populus adenopoda'' – Chinese aspen (China, south of ''P. tremula'') * ''Populus da ...

and

sallow Willows, also called sallows and osiers, of the genus ''Salix'', comprise around 350 species (plus numerous hybrids) of typically deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions. Most species are known ...

but has also been recorded on

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...

hazel Hazels are plants of the genus ''Corylus'' of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family, Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K ...

and poplar. The species overwinters as a

pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...

, sometimes remaining in this form for up to four years.

Subspecies

* ''Lomaspilis marginata amurensis'' (Hedemann, 1881) * ''Lomaspilis marginata marginata'' (Linnaeus, 1758) # ''The flight season refers to the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

. This may vary in other parts of the range.''

References

Chinery, Michael Michael Chinery (born 1938, in London) is an English naturalist. He studied in Cambridge where he graduated in natural sciences and anthropology. He edits '' Cecidology'', the journal of the British Plant Gall Society The British Plant Gall ...

(1986, reprinted 1991). ''Collins Guide to the Insects of Britain and Western Europe''. * Skinner, Bernard (1984). ''

The Colour Identification Guide to Moths of the British Isles ''The Colour Identification Guide to Moths of the British Isles (Macrolepidoptera)'' by Bernard Skinner is a single volume identification guide to the macromoths of Britain and Ireland. The first edition was published in 1984, and a second, revi ...

''.

External links

*
''Lepiforum e.V.''
Abraxini Moths described in 1758 Moths of Asia Moths of Europe Taxa named by Carl Linnaeus {{Abraxini-stub