The clitellum is a thickened

gland A gland is a Cell (biology), cell or an Organ (biology), organ in an animal's body that produces and secretes different substances that the organism needs, either into the bloodstream or into a body cavity or outer surface. A gland may also funct ...

ular and non-segmented section of the body wall near the head in

earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...

s and

leeches Leeches are segmented parasitic or predatory worms that comprise the subclass Hirudinea within the phylum Annelida. They are closely related to the oligochaetes, which include the earthworm, and like them have soft, muscular segmented bod ...

that secretes a viscid sac in which eggs are stored. It is located near the

anterior Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

end of the body, between the fourteenth and seventeenth segments. The number of the segments to where the clitellum begins and the number of segments that make up the clitellum are important for identifying earthworms. In microdrile earthworms, the clitellum has only one layer, resulting in a smaller quantity of eggs than that of the megadrile earthworms, which have larger multi-layered clitellum that have special cells that secrete

albumin Albumin is a family of globular proteins, the most common of which are the serum albumins. All of the proteins of the albumin family are water- soluble, moderately soluble in concentrated salt solutions, and experience heat denaturation. Alb ...

into the worms' egg sac. The clitellum is part of the reproductive system of clitellates, a subgroup of

annelid The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to vario ...

s which contains oligochaetes (earthworms) and hirudineans (leeches). The clitellum is a thick, saddle-like ring found in the epidermis (skin) of the worm, usually with a light-colored pigment. To form a cocoon for its eggs, the clitellum secretes a

viscous Viscosity is a measure of a fluid's rate-dependent resistance to a change in shape or to movement of its neighboring portions relative to one another. For liquids, it corresponds to the informal concept of ''thickness''; for example, syrup h ...

fluid. This organ is used in sexual reproduction of some annelids, such as leeches. In earthworms, the clitellum can only be seen when the worm is sexually mature. It may be white, orange-red or reddish-brown in colour. Earthworms are ready to mate when their clitellum is orange. In leeches, the clitellum appears during mating season, where it is used for both sexual reproduction and the secretion of a cocoon for the eggs. Its color is usually slightly lighter than that of the body of the annelid. Occasionally, living segments of the worm will be shed with the clitellum.

See also

References

Further reading