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''Clelandina'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of rubidgeine
gorgonopsia Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of Saber-toothed predator, sabre-toothed therapsids from the Middle Permian, Middle to the Upper Permian, roughly between 270 and 252 million years ago. ...
n from the
Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...
of ''Cistecephalus'' Assemblage Zone of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. It was first named by
Broom A broom (also known as a broomstick) is a cleaning tool, consisting of usually stiff fibers (often made of materials such as plastic, hair, or corn husks) attached to, and roughly parallel to, a cylindrical handle, the broomstick. It is thus a ...
in 1948. The type and only species is ''C. rubidgei''. It is relatively rare, with only four known specimens.


Description

''Clelandina rubidgei'' has an extraordinarily small
sclerotic ring The scleral ring or sclerotic ring is a hardened ring of plates, often derived from bone, that is found in the eyes of many animals in several groups of vertebrates. Some species of mammals, amphibians, and crocodilians lack scleral rings. The rin ...
relative to the size of its orbit, which implies that it was diurnal. It is the only rubidgeine with a preserved sclerotic ring, so it is unknown whether this trait was shared by other members of the subfamily. Like all rubidgeines, it was relatively large, with a skull up to 36 cm long. It had reduced dentition, with the teeth posterior to the canines being absent and replaced with a bony ridge. The skull has heavily pachyostosed, with massive rugose bosses.


Classification

''Clelandina'' shares many characteristics with the contemporary ''
Rubidgea ''Rubidgea'' is a genus of gorgonopsian from the upper Permian of South Africa and Tanzania, containing the species ''Rubidgea atrox''. The generic name ''Rubidgea'' is sometimes believed to be derived from the surname of renowned Karoo paleonto ...
'', and is currently recognized as the sister taxon of this genus. Together with '' Dinogorgon'' and '' Leontosaurus'', these genera form
Rubidgeini Rubidgeinae is an extinct subfamily of gorgonopsid therapsids known only from Africa. They were among the largest gorgonopsians, and their fossils are common in the Cistecephalus and Daptocephalus assemblage zones of the Karoo Basin. They l ...
, a clade of large gorgonopsians with distinctively robust, broad, and pachyostosed skulls. Below is a cladogram recovered by Christian Kammerer in 2016.


See also

*
List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ...


References


Further reading


paleodb.orgwww.paleofile.com Alphabetical list - C section.
*''The Origin and Evolution of Mammals'' by Tom Kemp. Published 2005, Oxford University Press. Gorgonopsia Prehistoric therapsid genera Wuchiapingian Permian synapsids of Africa Fossils of South Africa Fossil taxa described in 1948 Taxa named by Robert Broom {{paleo-therapsid-stub