The Cladonychiidae are a small family of
harvestman
The Opiliones (formerly Phalangida) are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, although the total number of ext ...
with about 33 described species, within the suborder
Laniatores.
Description
Members of this family range from less than two to about four millimeters in body length, with robust, spined
pedipalp
Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") a ...
s and rather short legs, although the second pair can be as long as two centimeter. Most Cladonychiidae are reddish brown to dark brown, but cave-dwelling species are pale yellow. Not all species have eyes.
[ (2007): Cladonychiidae Hadži, 1935. In: Pinto-da-Rocha ''et al.'' 2007: 179ff]
Distribution
The members of Cladonychiidae are found in Southern Europe and the United States.
[
]
Fossils
''Proholoscotolemon'' was recently found in Baltic amber
Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In M ...
.
Relationships
The genera ''Peltonychia
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* '' Peltonychia claviger ...
'', ''Holoscotolemon
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range fr ...
'', '' Erebomaster'', ''Theromaster
''Theromaster'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
'', ''Speleonychia
''Speleonychia'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
'', ''Briggsus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'', and
''Isolachus
''Isolachus'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from le ...
'' from the family Travuniidae
The Travuniidae are a small family of harvestman with little more than ten described species, within the suborder Laniatores.
Description
Travuniidae are at the most three millimeters long, with slender, unarmed legs and robust, strongly spined ...
and the former family Pentanychidae have been transferred to Cladonychiidae. This left Travuniidae with three genera from the Balkan
The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
region of Europe, ''Travunia
Travunia ( sh-Latn-Cyrl, separator=" / ", Travunija, Травунија; el, Τερβουνία, Tervounía; grc, Τερβουνία, Terbounía; la, Tribunia) was a South Slavic medieval principality that was part of Medieval Serbia (850–1 ...
'', '' Trojanella'', and '' Dinaria''.[
]
Name
The name of the former type genus ''Cladonychium'' (now synonymized with ''Erebomaster'') is derived from Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
"branched claw".[
]
Species
These genera and species belong to the family Cladonychiidae:[
* genus '']Arbasus
''Arbasus'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from less ...
'' Roewer, 1935
** species ''Arbasus caecus
''Arbasus'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from less ...
'' (Simon, 1911)
* genus ''Briggsus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'' Özdikmen & Demir, 2008
** species ''Briggsus bilobatus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'' (Briggs, 1971)
** species ''Briggsus clavatus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'' (Briggs, 1971)
** species ''Briggsus flavescens
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from les ...
'' (Briggs, 1971)
** species ''Briggsus hamatus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'' (Briggs, 1971)
** species ''Briggsus pacificus
''Briggsus'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about five described species in ''Briggsus'', found in the coastal forests of Oregon and Washington.
Species
These five species belong to the genus ''Briggsus ...
'' (Briggs, 1971)
* genus ''Buemarinoa
''Buemarinoa'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from l ...
'' Roewer, 1956
** species '' Buemarinoa patrizii'' Roewer, 1956
* genus '' Erebomaster'' Cope, 1872
** species '' Erebomaster acanthinus'' (Crosby & Bishop, 1924)
** species '' Erebomaster flavescens'' Cope, 1872
*** subspecies ''Erebomaster flavescens flavescens'' Cope, 1872
*** subspecies ''Erebomaster flavescens coecum'' (Packard, 1888) — Carter cave
** species '' Erebomaster weyerensis'' (Packard, 1888)
* genus ''Holoscotolemon
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range fr ...
'' Roewer, 1915
** species ''Holoscotolemon franzinii
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range fr ...
'' Tedeschi & Sciaky, 1994
** species ''Holoscotolemon jaqueti
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' (Corti, 1905)
** species '' Holoscotolemon lessiniense'' Martens, 1978
** species ''Holoscotolemon lessiniensis
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' Martens, 1978
** species ''Holoscotolemon monzinii
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' Tedeschi & Sciaky, 1994
** species ''Holoscotolemon naturae
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' Tedeschi & Sciaky, 1994
** species ''Holoscotolemon oreophilus
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' Martens, 1978
** species ''Holoscotolemon querilhaci
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' (Lucas, 1864)
** species ''Holoscotolemon unicolor
''Holoscotolemon'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about eight described species in ''Holoscotolemon'', found in Europe.
Species
These eight species belong to the genus ''Holoscotolemon'':
* ''Holoscotol ...
'' Roewer, 1915
* genus ''Isolachus
''Isolachus'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from le ...
'' Briggs, 1971
** species '' Isolachus spinosus'' Briggs, 1971
* genus ''Peltonychia
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* '' Peltonychia claviger ...
'' Roewer, 1935
** species ''Peltonychia clavigera
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* '' Peltonychia claviger ...
'' (Simon, 1879)
** species ''Peltonychia gabria
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' Roewer, 1935
** species ''Peltonychia leprieuri
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' (Lucas, 1861)
** species ''Peltonychia leprieurii
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
''
** species ''Peltonychia navarica
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' (Simon, 1879)
** species ''Peltonychia piochardi
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' (Simon, 1872)
** species ''Peltonychia postumicola
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' (Roewer, 1935)
** species ''Peltonychia sarea
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' (Roewer, 1935)
** species ''Peltonychia tenuis
''Peltonychia'' is a genus of armoured harvestmen in the family Cladonychiidae. There are about nine described species in ''Peltonychia'', found in Europe.
Species
These nine species belong to the genus ''Peltonychia'':
* ''Peltonychia clavigera ...
'' Roewer, 1935
* genus ''Speleonychia
''Speleonychia'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
'' Briggs, 1974
** species '' Speleonychia sengeri'' Briggs, 1974
* genus ''Theromaster
''Theromaster'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
'' Briggs, 1969
** species ''Theromaster archeri
''Theromaster'' is a genus of armoured harvestmen in the family Cladonychiidae
The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.
Description
Members of this family range from ...
'' (Goodnight & Goodnight, 1942)
** species ''Theromaster brunneus
''Theromaster brunneus'' is a species of armoured harvestman in the family Travuniidae
The Travuniidae are a small family of harvestman with little more than ten described species, within the suborder Laniatores.
Description
Travuniidae ar ...
'' (Banks, 1902)
* genus † '' Proholoscotolemon'' Ubick & Dunlop, 2005
** species † '' Proholoscotolemon nemastomoides'' (Koch & Berendt, 1854)
Footnotes
References
* (eds.) (2007): Harvestmen - The Biology of Opiliones. ''Harvard University Press''
{{Taxonbar, from=Q2974943
Harvestman families
Extant Eocene first appearances
Harvestmen