Cistus Chinamadensis
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''Cistus chinamadensis'' is a
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
by
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the family
Cistaceae The Cistaceae are a small family of plants (rock-rose or rock rose family) known for their beautiful shrubs, which are profusely covered by flowers at the time of blossom. This family consists of about 170(-200) species in eight genera that are ...
, with purple-pink flowers, first described in 1991. It is endemic to the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
, where three subspecies occur on three separate islands (
Tenerife Tenerife ( ; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands, an Autonomous communities of Spain, autonomous community of Spain. With a land area of and a population of 965,575 inhabitants as of A ...
,
La Gomera La Gomera () is one of Spain's Canary Islands, in the Atlantic Ocean off the coast of Africa. With an area of , it is the third-smallest of the archipelago's eight main islands. It belongs to the province of Santa Cruz de Tenerife. La Gomer ...
and
El Hierro El Hierro (), nicknamed ''Isla del Meridiano'' (the "Meridian Island"), is the farthest south and west of the Canary Islands (an autonomous community of Spain), in the Atlantic Ocean off the coast of Africa, with a population of 11,659 (2023). ...
). The species has been assessed as endangered in the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
, being known only from small separated areas and facing a variety of threats.


Description

''Cistus chinamadensis'' is a
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
, usually tall. The woody stems have dark brown bark that easily frays and strips off. The upper branches have a dense velvety covering of fine hairs (
indumentum In biology, an indumentum (Latin, literally: "garment") is a covering of trichomes (fine "hairs") on a plant or of bristles (rarely scales) of an insect. Plants The indumentum on plants can have a wide variety of functions, including as ...
), beige to off-white in colour. The oppositely arranged leaves are light green (in subsp. ''gomerae'') or greyish green (in subsp. ''chinamadensis''), around long by wide, with a pointed tip. The leaves have three prominent veins. Opposite pairs of leaves are joined at the base by a long sheath, the outside of which is furrowed and the inside covered in hairs (as is the outside in subsp. ''gomerae''). The upper sides of the leaves are more-or-less smooth, the lower sides rough and reticulate, to a varying degree between the subspecies. The flowers are arranged in open, slightly branched
cymes In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis ( ped ...
, with 4–8 flowers to each
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
. The
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s have a short tooth, up to long, at the apex and are of two distinct sizes. Two outer sepals are around long by wide; three inner sepals are considerably larger, around long by wide. The
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s are pink with yellowish bases, more-or-less rounded, about long and wide. The yellow
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s are somewhat shorter than the sepals. The
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
is high, topped by a
style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...
about long. In its native habitat, ''C. chinamadenis'' flowers in May and produces seeds in June to July.


Taxonomy and phylogeny

''Cistus chinamadensis'' was first described in 1991 by Ángel Bañares Baudet and Pedro Romero Manrique. The species was first found at Roque de los Pinos near to the village of Chinamada in the
Anaga Anaga massif () is a mountain range in the northeastern part of the island of Tenerife in the Canary Islands. The highest point is 1,024 m (Cruz de Taborno). It stretches from the Punta de Anaga in the northeast to Cruz del Carmen in the sou ...
region of
Tenerife Tenerife ( ; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands, an Autonomous communities of Spain, autonomous community of Spain. With a land area of and a population of 965,575 inhabitants as of A ...
; hence the specific epithet ''chinamadensis'' meaning "from Chinamada". Two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
were initially described, ''C. chinamadensis'' subsp. ''chinamadensis'' and ''C. ch.'' subsp. ''gomerae''. In 2005 a further subspecies, ''C. ch.'' subsp. ''ombriosus'', was described by Jean-Pierre Demoly and M. Marrero. A 2011 molecular phylogenetic study placed ''C. chinamadensis'' as a member of the purple and pink flowered clade of ''Cistus'' species, along with some other Canary Island endemics (''
Cistus asper ''Cistus asper'' is a shrubby species of flowering plant in the family Cistaceae with purple-pink flowers. It was first described in 2005 and is endemic to El Hierro in the Canary Islands The Canary Islands (; ) or Canaries are an archip ...
'', '' Cistus horrens'', '' Cistus ocreatus'', and '' Cistus symphytifolius''), although the three subspecies did not form a clade, with some analyses separating ''C. ch.'' subsp. ''ombriosus'' in particular from the other two subspecies.


Distribution and habitat

''Cistus chinamadensis'' is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
. The three subspecies are found on different islands: ''C. ch.'' subsp. ''chinamadensis'' in the north of
Tenerife Tenerife ( ; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands, an Autonomous communities of Spain, autonomous community of Spain. With a land area of and a population of 965,575 inhabitants as of A ...
in three locations in the
Anaga Anaga massif () is a mountain range in the northeastern part of the island of Tenerife in the Canary Islands. The highest point is 1,024 m (Cruz de Taborno). It stretches from the Punta de Anaga in the northeast to Cruz del Carmen in the sou ...
region at altitudes of around ; ''C. ch.'' subsp. ''gomerae'' in one natural location and several introduced locations in
La Gomera La Gomera () is one of Spain's Canary Islands, in the Atlantic Ocean off the coast of Africa. With an area of , it is the third-smallest of the archipelago's eight main islands. It belongs to the province of Santa Cruz de Tenerife. La Gomer ...
at around ; and ''C. ch.'' subsp. ''ombriosus'' in one location in
El Hierro El Hierro (), nicknamed ''Isla del Meridiano'' (the "Meridian Island"), is the farthest south and west of the Canary Islands (an autonomous community of Spain), in the Atlantic Ocean off the coast of Africa, with a population of 11,659 (2023). ...
at .


Conservation

In 2011 when its status was assessed as "endangered" according to the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
criteria, ''Cistus chinamadensis'' was known from seven locations in the Canary Islands. ''C. ch.'' ssp. ''chinamadensis'' from Tenerife was present in three locations; its population appeared to be stable or increasing. ''C. ch.'' ssp. ''gomerae'' from La Gomera was found in two natural locations and has since been introduced to four others. Its range appeared to be decreasing. ''C. ch.'' subsp. ''ombriosus'' from El Hierro was present in a single location with an area of about . Threats to the species include a variety of natural hazards, such as landslides, fires and drought, and biological hazards, such as grazing.


References


External links

* {{Taxonbar, from=Q15330461 chinamadensis Plants described in 1991 Endemic flora of the Canary Islands