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''Cirsium scariosum'' is a species of
thistle Thistle is the common name of a group of flowering plants characterised by leaves with sharp prickles on the margins, mostly in the family Asteraceae. Prickles can also occur all over the planton the stem and on the flat parts of the leaves ...
known by the common names meadow thistle, elk thistle and dwarf thistle. It is native to much of western North America from
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
and
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost Provinces and territories of Canada, province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include ...
, south to
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
. There are also isolated populations on the Canadian Atlantic Coast, on the
Mingan Archipelago The Mingan Archipelago is an archipelago located east of Quebec, Canada. It consists of a chain of about 40 islands. Starting but 124 miles from the end of the road along the north shore of the St. Lawrence River (Le Fleuve), the Mingan Arc ...
in
Québec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirteen p ...
,Flora of North America, Meadow or elk thistle, chardon écailleux, ''Cirsium scariosum'' Nuttall
/ref> where it is called the ''Mingan thistle''. It was first published in Trans. Amer. Philos. Soc., ser.2, vol.7 on page 420 in 1841.


Description

''Cirsium scariosum'' is a variable species growing in a variety of habitat types. It is made up of several geographic races, most of which have been previously classified as species. The races
intergrade In zoology, intergradation is the way in which two distinct subspecies are connected via areas where populations are found that have the characteristics of both. There are two types of intergradation: primary and secondary intergradation. Primar ...
but their morphologies can also be quite different. This is generally a
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...
or
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
herb. It takes three main forms, a stemless, flat rosette with a cluster of
flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
in the centre, a mounding form with a short, erect stem, or a fully erect form reaching up to in height. When there is a stem it is usually fleshy, ridged, and woolly in texture. The leaves are sharply toothed or cut into toothed lobes, lined with spines, and up to at their longest near the base of the plant. The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
holds several
flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
, each up to 4 centimeters long and 5 cm wide. The
flower head A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
is lined with
phyllaries In botanical terminology, a phyllary, also known an involucral bract or tegule, is a single bract of the involucre of a composite flower. The involucre is the grouping of bracts together. Phyllaries are reduced leaf-like structures that form one ...
which may have spines and teeth and filled with white to purple
disc floret The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae we ...
s but no
ray florets The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae we ...
. The fruit is a compressed
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not op ...
a few millimetres long topped with a pappus which may be 3 centimeters in length.


Varieties

As accepted by
Flora of North America The ''Flora of North America North of Mexico'' (usually referred to as ''FNA'') is a multivolume work describing the native plants and naturalized plants of North America North America is a continent in the Northern Hemisphere and a ...
; *''Cirsium scariosum'' var. ''americanum'' -
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
,
Colorado Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the wes ...
,
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and W ...
,
Nevada Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
,
Oregon Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idah ...
,
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
,
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the sou ...
, Baja California *''Cirsium scariosum'' var. ''citrinum'' - California, Baja California *''Cirsium scariosum'' var. ''coloradense'' -
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a U.S. state, state in the Southwestern United States. It is the list of U.S. states and territories by area, 6th largest and the list of U.S. states and territories by population, 14 ...
, Colorado,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
, Utah,
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the sou ...
*''Cirsium scariosum'' var. ''congdonii'' - California,
Nevada Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
*''Cirsium scariosum'' var. ''robustum'' - California, Oregon *''Cirsium scariosum'' var. ''scariosum'' - Quebec, Alberta, British Columbia, California, Colorado, Idaho,
Montana Montana () is a U.S. state, state in the Mountain states, Mountain West List of regions of the United States#Census Bureau-designated regions and divisions, division of the Western United States. It is bordered by Idaho to the west, North ...
, Oregon, Utah,
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
, Wyoming *''Cirsium scariosum'' var. ''thorneae'' - Utah, Nevada, Idaho, Colorado *''Cirsium scariosum'' var. ''toiyabense''- Nevada, Oregon, Idaho


References


External links


Jepson Manual TreatmentCalphotos Photo gallery, University of California
scariosum Plants described in 1841 Flora of North America Taxa named by Thomas Nuttall {{Cynareae-stub