Cirrhitidae, the hawkfishes, are a
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of marine perciform
ray-finned fishes found in tropical seas and which are associated with coral reefs.
Taxonomy
The Cirrhitidae were first recognised as a family by the
Scots
Scots usually refers to something of, from, or related to Scotland, including:
* Scots language, a language of the West Germanic language family native to Scotland
* Scots people, a nation and ethnic group native to Scotland
* Scoti, a Latin na ...
-born
Australian
Australian(s) may refer to:
Australia
* Australia, a country
* Australians, citizens of the Commonwealth of Australia
** European Australians
** Anglo-Celtic Australians, Australians descended principally from British colonists
** Aboriginal ...
naturalist William Sharp Macleay
William Sharp Macleay or McLeay (21 July 1792 – 26 January 1865) was a British civil servant and entomologist. He was a prominent promoter of the Quinarian system of classification.
After graduating, he worked for the British embassy in Pari ...
in 1841.
[ It is one of the 5 constituent families in the ]superfamily
SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Cirrhitoidea which is classified in the suborder
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
Percoidei
Percoidei is one of 3 suborders of bony fishes in the order Perciformes. Many commercially harvested fish species are considered to be contained in this suborder, including the snappers, groupers, basses, goatfishes and perches.
Divisions
Th ...
of the order Perciformes
Perciformes (), also called the Percomorpha or Acanthopteri, is an order or superorder of ray-finned fish. If considered a single order, they are the most numerous order of vertebrates, containing about 41% of all bony fish. Perciformes means ...
. Within the Cirrhitoidea, the Cirrhitidae is probably the most basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
family. They have been placed in the order Centrarchiformes by some authorities, as part of the superfamily Cirrhitoidea, however, the 5th edition of Fishes of the World
''Fishes of the World'' by the American Ichthyology, ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classificat ...
does not recognise the Centrarchiformes.[ The name of the family is taken from that of the ]genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Cirrhitus'' which is derived from ''cirrhus'' meaning a "lock of hair" or "a barbel", thought to be a reference to lower, unbranched rays of the pectoral fins which Bernard Germain de Lacépède
Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's great work, the ...
termed as “''barbillons''”, which means "barbels" in his description
Description is the pattern of narrative development that aims to make vivid a place, object, character, or group. Description is one of four rhetorical modes (also known as ''modes of discourse''), along with exposition, argumentation, and na ...
of the type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
of the genus '' C. maculatus'', and which he thought to be “false” pectoral fins
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
. Another possibility is that the name refers to cirri
Giovanni Battista Cirri (1 October 1724 – 11 June 1808) was an Italian cellist and composer in the 18th century.
Biography
Cirri was born in Forlì in the Emilia-Romagna Region of Italy. He had his first musical training with his brother ...
extending from the tips of the spines in the dorsal fin
A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
spines, although Lacépède did not mention this feature.
Genera
The following 12 genera are classified within the Cirrhitidae, containing a total of 33 species:
* ''Amblycirrhitus
''Amblycirrhitus'' is a genus of ray-finned fishes, hawkfishes belonging to the family Cirrhitidae. These fishes are found on tropical reefs worldwide.
Taxonomy
''Amblycirrhitus'' was originally described as a genus in 1862 by the American ic ...
'' Gill
A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they ar ...
, 1862
* ''Cirrhitichthys
''Cirrhitichthys'' is a genus of marine ray-finned fish, hawkfishes, from the family Cirrhitidae. They are found on tropical reefs in the Indian and western Pacific oceans. Some species can be found in the aquarium trade.
Taxonomy
''Cirrhitich ...
'' Bleeker, 1857
* ''Cirrhitops
''Cirrhitops'' is a genus of marine ray-finned fish, hawkfishes from the family Cirrhitidae. They are found on tropical reefs of the Indian Ocean and the Pacific Ocean.
Taxonomy
''Cirrhitops'' was described as a genus in 1951 by the South Afric ...
'' J.L.B Smith, 1951
* ''Cirrhitus
''Cirrhitus'' is a genus of marine ray-finned fish, hawkfishes from the family Cirrhitidae. The species in this genus are found on tropical reefs worldwide.
Taxonomy
''Cirrhites'' was first formally described as a genus in 1803 by the French ...
'' Lacepède, 1803
* ''Cristacirrhitus
The blackspotted hawkfish (''Cristacirrhitus punctatus'') is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is native to rocky shores of the western Indian Ocean. This species grows to in total length. ...
'' Randall Randall may refer to the following:
Places
United States
*Randall, California, former name of White Hall, California, an unincorporated community
* Randall, Indiana, a former town
*Randall, Iowa, a city
*Randall, Kansas, a city
*Randall, Minnesot ...
, 2001
* '' Cyprinocirrhites'' Tanaka, 1917
* '' Isocirrhitus'' Randall, 1963
* ''Itycirrhitus
Wilhelm's hawkfish (''Itycirrhitus wilhelmi'') is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is found in tropical waters at depths of over rocky substrates. It is only known around Pitcairn Island ...
'' Randall 2001
* ''Neocirrhites
The flame hawkfish (''Neocirrhites armatus'') is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is native to tropical reefs of the Pacific Ocean at depths of from . This species is also found in the aqua ...
'' Castelnau, 1873
* '' Notocirrhitus'' Randall, 2001
* '' Oxycirrhites'' Bleeker, 1857
* ''Paracirrhites
''Paracirrhites'' is a genus of marine ray-finned fish, hawkfishes belonging to the family Cirrhitidae. These fishes are found on tropical reefs of the Indian Ocean and Pacific Ocean.
Taxonomy
''Paracirrhites'' was described in 1874 by the Dut ...
'' Bleeker, 1874
Characteristics
Cirrhitidae hawkfishes are roughly oblong in shape[ with a body which has a depth which is 21% to 50% of its ]standard length
Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.
Overall length
* Standard length (SL) is the length of a fish ...
. They have a fringe of cirri on the rear edge of the forward nostrils. There are two poorly developed spines, on the gill cover. The outer row of teeth on the jaws are canine-like, the longest normally being located at the front of the upper jaw and the middle of the lower jaw. Inside this row, there is a band of bristle-like teeth, wider in the front. The dorsal fin
A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
is continuous, having 10 spines and 11-17 soft rays, it has an incision separating the spiny and soft-rayed parts. The anal fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
contains 3 spines and 5–7, typically 6, soft rays. There are 14 pectoral fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
rays with the lowest 5-7 rays unbranched and normally thickened, with deep notches in the membranes separating these lower rays. There is a single spine in the pelvic fin
Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods.
Structure and function Structure
In actinopterygians, the pelvic fin consists of two e ...
s as well as 5 soft rays. The scales
Scale or scales may refer to:
Mathematics
* Scale (descriptive set theory), an object defined on a set of points
* Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original
* Scale factor, a number ...
are cycloid
In geometry, a cycloid is the curve traced by a point on a circle as it rolls along a straight line without slipping. A cycloid is a specific form of trochoid and is an example of a roulette, a curve generated by a curve rolling on another ...
and they lack a swimbladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...
. The colour and pattern vary between species. The maximum length attained is around ,[ although around is more typical.][ Most species are quite small and colourfully patterned.][
]
Distribution and habitat
Cirrhitidae hawkfishes are found in the tropical western and eastern Atlantic, Indian and Pacific, mainly in the Indo-West Pacific
The Indo-Pacific is a vast biogeographic region of Earth.
In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region.[ They are ]benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
fishes which are found on coral reefs or rocky substrates, mostly inhabiting shallow water.[
]
Biology
Cirrhitidae fishes use their robust lower pectoral-fin rays to wedge into position where they will be subjected to the forces of currents and waves. They are carnivorous fishes, their main prey being benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
crustaceans. One species, '' Cyprinocirrhitus polyactis'', mainly feeds on zooplankton, although it is frequently encountered resting on the substrate.[ Hawkfish frequently sit and wait on the higher parts of their habitat, diving onto prey items seen underneath them, in a similar manner to some ]hawk
Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica.
* The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfa ...
species, hence the name hawkfish.
Fisheries and utilisation
Cirrhitidae hawkfishes are mostly too small to be of interest to fisheries. The 3 largest species are occasionally fished for as food fish
Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is inges ...
. A few of the smaller more colourful species, particularly ''Neocirrhites armatus
The flame hawkfish (''Neocirrhites armatus'') is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is native to tropical reefs of the Pacific Ocean at depths of from . This species is also found in the aqua ...
'' and ''Oxycirrhites typus
The longnose hawkfish (''Oxycirrhites typus'') is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is found on tropical reefs of the Indian Ocean and the Pacific Ocean, where it can be found at depths around ...
'', are collected for the aquarium trade.[
]
Gallery
Image:Dwarf HawkFish.jpg, Dwarf hawkfish (''Cirrhitichthys falco''), Sipadan
Sipadan ( ms, Pulau Sipadan) is the only oceanic island in Malaysia, rising from the seabed. It is located in the Celebes Sea off the east coast of Sabah, Malaysia. It was formed by living corals growing on top of an extinct volcanic cone t ...
, Malaysia
Image:Long-nose_Hawkfish.jpg, Longnose hawkfish (''Oxycirrhites typus''), Galápagos Islands
The Galápagos Islands ( es, Islas Galápagos) are an archipelago of volcanic islands in the Eastern Pacific, located around the Equator west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with ...
Image:Coral_Hawkfish.jpg, Coral hawkfish
The coral hawkfish (''Cirrhitichthys oxycephalus''), the pixy hawkfish or sharp-headed hawkfish, is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is native to tropical reefs of the Indian Ocean and ...
(''Cirrhitichthys oxycephalus''), Galápagos Islands
Image:Arc-Eye Hawkfish.jpg, Arc-eye hawkfish
The arc-eye hawkfish (''Paracirrhites arcatus''), the ringeye hawkfish, horseshoe hawkfish or whiteline hawkfish, is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is found in shallow waters in the tropic ...
(''Paracirrhites arcatus''), Kona District, Hawaii
Kona is a ''moku'' or district on the Big Island of Hawaii in the State of Hawaii, known for its Kona coffee and the location of the Ironman World Championship Triathlon. In the current system of administration of Hawaii County, the ''moku'' ...
Image:Cirrhitichthys aprinus.JPG, Spotted hawkfish (''Cirrhitichthys aprinus''), Lembeh Straits, Indonesia
Image:Coral Hawkfish .jpg, Coral hawkfish
The coral hawkfish (''Cirrhitichthys oxycephalus''), the pixy hawkfish or sharp-headed hawkfish, is a species of marine ray-finned fish, a hawkfish belonging to the family Cirrhitidae. It is native to tropical reefs of the Indian Ocean and ...
(''Cirrhitichthys oxycephalus''), Great barrier reef, Australia
References
External links
*
{{Taxonbar, from=Q265485
Fish of Hawaii
Ray-finned fish families
Cirrhitoidea