''Choconsaurus'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of herbivorous

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

belonging to the group Titanosauriformes, which lived in the area of present-day

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...

at the end of the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

Discovery and naming

The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

(specimen MMCh-PV 44/10) was found in Villa El Chocón in Neuquén province,

by Viviana Moro before 1996. Subsequent field campaigns carried out by the Ernesto Bachmann Paleontological Museum and the Museum of Geology and Paleontology of the National University of Comahue between 1996 and 2002 discovered more remains pertaining to ''Choconsaurus'', with several of the specimens described by Calvo (1999) under "

Titanosauridae Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of '' Malawisaurus'' and '' Saltasaurus ...

indet.".Calvo, J.O. (1999). Dinosaurs and other vertebrates of the Lake Ezequiel Ramos Mexía Area, Neuquén, Patagonia, Argentina. ''National Museum Monographs'' 15: 13–45. In 2017, the species type ''Choconsaurus baileywillisi'' was named and described by Edith Simón, Leonardo Salgado and Jorge Orlando Calvo. The genus name refers to its discovery in El Chocón. For its part, the name of the species is in homage to the American geologist Bailey Willis, who traced the

stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers ( strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...

of the area between 1910 and 1914.

Description

The

specimen was found in a layer of the Huincul Formation dating to the end of the Cenomanian epoch. It consists of a partial skeleton lacking a skull. The vertebrae of the neck, back and tail and parts of the limbs were preserved. This constitutes the most complete skeleton known of a basal titanosaur in 2017. The describing authors have identified some distinctive features. In the

cervical vertebra In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...

e, the upper edge of the face of the posterior joint was hardly developed. In the first vertebrae there are very large secondary protuberances on both sides of the hyposphene. The central and posterior vertebrae have an additional crest located between the posterior crest that extends from the lateral protrusion to the vertebral body and the secondary crest running parallel to the main crest. The frontal vertebrae of the tail have a protrusion in the posterior hyposphenic secondary joint.

Classification

''Choconsaurus'' was classified in its description article of 2017 within the clade Titanosauria in a

basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...

position, outside the clade Eutitanosauria.

References

{{Taxonbar, from=Q41508018 Titanosaurs Cenomanian life Late Cretaceous dinosaurs of South America Cretaceous Argentina Fossils of Argentina Huincul Formation Fossil taxa described in 2017

Discovery and naming

Description

Classification

See also

References