Chiriqui Foliage-gleaner
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The Chiriqui foliage-gleaner (''Automolus exsertus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is found in
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
and
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
.


Taxonomy and systematics

The Chiriqui foliage-gleaner was originally described as a species but soon was reconsidered as a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of what is now the
ochre-throated foliage-gleaner The ochre-throated foliage-gleaner (''Automolus ochrolaemus'') is a species of bird in the Furnariinae subfamily of the ovenbird family Furnariidae. It is found in Panama and every mainland South American country except Argentina, Chile, Paragu ...
(''A. ochrolaemus'').Remsen, Jr., J. V. and H. F. Greeney (2020). Chiriqui Foliage-gleaner (''Automolus exsertus''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.butfog4.01 retrieved September 7, 2023 A 2017 publication detailed significant differences in its plumage and vocalizations from the other ochre-throated subspecies. Based on that data, the North American Classification Committee of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
, the
International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...
, and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
elevated it to a full species.Chesser, R. T., S. M. Billerman, K. J. Burns, C. Cicero, J. L. Dunn, B. E. Hern ndez-Ba os, R. A. Jim nez, A. W. Kratter, N. A. Mason, P. C. Rasmussen, J. V. Remsen, Jr., and K. Winker. 2023. Check-list of North American Birds (online). American Ornithological Society. https://checklist.americanornithology.org/taxa/Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022 As of late 2023,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' (HBW) still treats the taxon as a subspecies of ''A. ochrolaemus'', which it calls the buff-throated foliage-gleaner.HBW and BirdLife International (2022) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 7. Available at: https://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v7_Dec22.zip retrieved December 13, 2022 The Chiriqui foliage-gleaner is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
.


Description

The Chiriqui foliage-gleaner is long and weighs . It is a fairly large member of its genus and has a heavy bill. The sexes have the same plumage. Adults have a mostly dark brownish face with a bold buff eyering, faint reddish streaks on the ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
, and a slightly buffy malar area. Their crown and nape are dark brown with a faint blackish brown scallop pattern. Their back and rump are rich dark brown that blends to dark chestnut uppertail coverts. Their wing coverts are rich dark brown and their flight feathers slightly paler and more rufescent. Their tail is dark chestnut. Their throat is ochraceous-buff, their breast is streaked with medium brown and ochraceous buff that fades in its lower part, and their belly is brown. Their flanks are a darker and more rufescent brown and their undertail coverts bright rufous. Their iris is brown to dark brown, their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
blackish horn to horn-brown, their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
horn to gray, and their legs and feet olive, greenish brown, or greenish gray.


Distribution and habitat

The Chiriqui foliage-gleaner is found on the Pacific slope from Costa Rica south of the dry northwest into western Panama as far as
Veraguas Province Veraguas () is a province of Panama, located in the centre-west of the country. The capital is the city of Santiago de Veraguas. It is the only Panamanian province to border both the Atlantic and Pacific oceans. It covers an area of 10,587.6&n ...
. It inhabits tropical
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, live oak, acacias, magnolia, and banksia in more temperate zo ...
, mature
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
, and coffee plantations from sea level to .R. Terry Chesser, Kevin J. Burns, Carla Cicero, Jon L. Dunn, Andrew W. Kratter, Irby J. Lovette, Pamela C. Rasmussen, J. V. Remsen, Jr., Douglas F. Stotz, Benjamin M. Winger, and Kevin Winker. "Fifty-ninth supplement to the American Ornithological Society s ''Check-list of North American Birds''". ''The Auk'' 2018, vol. 135:798-813 retrieved December 13, 2022


Behavior


Movement

The Chiriqui foliage-gleaner is a year-round resident throughout its range.


Feeding

The Chiriqui foliage-gleaner feeds mostly on a variety of insects, spiders, and vertebrates like small lizards. It forages singly or in pairs and often (perhaps usually) joins
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. It forages from the forest's dense undergrowth to its lower mid-storey, acrobatically gleaning and pulling prey from
epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
s, debris, and especially from clumps of dead leaves as it hops along branches and vines. It has also been observed foraging on the ground by flipping aside leaf litter.


Breeding

The Chiriqui foliage-gleaner's breeding season spans from February to May. It is monogamous. It excavates a tunnel up to about long in an earthen bank and constructs a shallow cup nest of leaf rachides in a chamber at its end. The clutch size is two or three eggs. The incubation period is 20 to 21 days and fledging occurs about 18 days after hatch. Both parents incubate the clutch and provision the nestlings.Skutch, A. F. (1969). Life histories of Central American birds III. Pacific Coast Avifauna 35:1–580.


Vocalization

The Chiriqui foliage-gleaner's song is "a loud, harsh rattle lasting 2–5 seconds, repeated 10–30 times per minute". Its call is "a nasal ''rack'' or hard ''tchehrr''.


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
follows
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's taxonomy and so has assessed the pre-split buff-throated foliage gleaner as a whole, with no separate evaluation of the Chiriqui foliage-gleaner. The species is considered common to fairly common and appears to tolerate some forest fragmentation.


References

{{Taxonbar, from=Q60793238 Chiriqui foliage-gleaner Birds of Central America Chiriqui foliage-gleaner Taxobox binomials not recognized by IUCN