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''Chimaphila umbellata'', the umbellate wintergreen, pipsissewa, or prince's pine, is a small perennial flowering plant found in dry woodlands, or sandy soils. It is native throughout the cool temperate
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. It grows 10–35 cm tall, and has evergreen shiny, bright green, toothed leaves arranged in opposite pairs or whorls of 3–4 along the stem. Leaves have a shallowly toothed margin, where the teeth have fine hairs at their ends. The flowers are white or pink, produced in a small umbel of 4–8 together.


Ecology

Although it has green leaves year-round, it receives a significant portion of its nutrition from fungi in the soil (that is, it is a partial myco-heterotroph, which is not surprising as related plants, such as ''
Pyrola ''Pyrola'' is a genus of evergreen herbaceous plants in the family Ericaceae. Under the old Cronquist system it was placed in its own family Pyrolaceae, but genetic research showed it belonged in the family Ericaceae. The species are commonly k ...
'', are partial or full myco-heterotrophs).


Taxonomy

There are four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: *''Chimaphila umbellata'' subsp. ''umbellata'' – Europe, Asia *''Chimaphila umbellata'' subsp. ''acuta'' – southwestern North America *''Chimaphila umbellata'' subsp. ''cisatlantica'' – northeastern North America *''Chimaphila umbellata'' subsp. ''occidentalis'' – northwestern North America


Uses

Some Plateau Indian tribes used a boil of prince's pine to treat tuberculosis. The twentieth century Appalachian folk healer Clarence "Catfish" Gray, "Man of the Woods", credited pipsissewa with curing his own heart problems and included it in his 15 herb cure-all "bitters." It is used as a flavoring in candy and soft drinks, particularly root beer. The roots and leaves of ''Chimaphila umbellata'' can be boiled to create tea. Recent investigations show the anti-proliferative effect of ''Chimaphila umbellata'' in human breast cancer cells (MCF-7). Das, N., Samantaray, S., Ghosh, C., Kushwaha, K., Sircar, D. and Roy, P., 2021. ''Chimaphila umbellata'' extract exerts anti-proliferative effect on human breast cancer cells via RIP1K/RIP3K-mediated necroptosis. Phytomedicine Plus, p.100159.


Name

"Pipsissewa" is a
Cree The Cree ( cr, néhinaw, script=Latn, , etc.; french: link=no, Cri) are a Indigenous peoples of the Americas, North American Indigenous people. They live primarily in Canada, where they form one of the country's largest First Nations in Canada ...
name meaning "It-breaks-into-small-pieces".


References

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External links


Flora Europaea: ''Chimaphila umbellata'' distributionlinnaeus.nrm.se: range map with the different subspecies of the ''Chimaphila umbellata''-complex Jepson Manual treatment – ''Chimaphila umbellata''USDA Plants Profile: Chimaphila umbellata
{{Taxonbar, from=Q158939 umbellata Flora of North America Flora of Europe Flora of temperate Asia Plants used in traditional Native American medicine Medicinal plants of Asia Medicinal plants of Europe Plants described in 1753 Rhizomatous plants Stoloniferous plants Taxa named by Carl Linnaeus