Chilantaisaurus
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''Chilantaisaurus'' (" lizard") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of large
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
, possibly a
neovenatorid Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
or a primitive
coelurosaur Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
, from the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Ulansuhai Formation The Ulansuhai Formation () is a geological Formation (geology), formation in Inner Mongolia, north China. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
(
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
age, about 92 million years ago). The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''C. tashuikouensis'', was described by Hu in 1964.


Description

''Chilantaisaurus'' was a large theropod, measuring long and weighing . While Brusatte ''et al.'' (2010) estimated that ''Chilantaisaurus'' might have weighed about based on femur length similar to that of ''
Tyrannosaurus ''Tyrannosaurus'' () is a genus of large theropod dinosaur. The type species ''Tyrannosaurus rex'' ( meaning 'king' in Latin), often shortened to ''T. rex'' or colloquially t-rex, is one of the best represented theropods. It lived througho ...
'', Persons ''et al.'' (2020) argued that greater femoral circumference indicates the greater capacity to withstand greater locomotor loads, not greater body mass.


Classification

Hu considered ''Chilantaisaurus'' to be a
carnosaur Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
related to ''
Allosaurus ''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
'', though some subsequent studies suggested that it may be a spinosauroid, possibly a primitive member of the
spinosaurid Spinosauridae (or spinosaurids) is a clade or Family (taxonomy), family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. ...
family (Sereno, 1998; Chure, 2000; Rauhut, 2001) because it had large claws on the forelimbs thought to be unique to that group. Other studies suggested that it could be a member of an alternate offshoot of neotetanuran theropods, with some similarities to allosauroids, spinosauroids, and
coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
ns. A 2009 study noted that it was difficult to rule out the possibility that ''Chilantaisaurus'' was the same animal as the carnosaur ''
Shaochilong ''Shaochilong'' (meaning "shark toothed dragon") is an extinct genus of tetanuran theropod dinosaurs from the Early Cretaceous (Barremian to Albian stages) Miaogou Formation of China, though a more restrictive suggestion from the Aptian to the A ...
'', which was thought to be from the same geological formation (a later study indicated that the latter genus actually derives from the Early Cretaceous
Miaogou Formation The Miaogou Formation is a geological formation in Inner Mongolia, north China. While its absolute age is uncertain, it has been estimated to represent Early Cretaceous sediments based on the faunal composition. Dinosaur remains are among the fos ...
). However, they did note an enormous size difference between the two. Further study by Benson, Carrano and Brusatte found that it was not as closely related to ''Shaochilong'' as first thought, but that it was a carnosaur (of the family
Neovenatoridae Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
), closely related to ''Allosaurus'' as Hu had initially thought. Phylogenetic analysis published by Porfiri et al. in 2018 recovered ''Chilantaisaurus'' as a primitive
coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
n. Several species have been described based on very poor remains. The species ''"Chilantaisaurus" sibiricus'' (previously informally known as either ''
Allosaurus ''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
? sibiricus'' or ''
Antrodemus ''Antrodemus'' ("chamber bodied") is a dubious genus of theropod dinosaur from the Upper Jurassic, probably the Morrison Formation, of Middle Park, Colorado. It contains one species, ''Antrodemus valens'', first described and named as a species ...
? sibiricus'') is based on a single distal
metatarsal The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
discovered in 1915 in the
Turginskaya Svita The Turginskaya Svita is an Early Cretaceous period geologic formation located in Russia. Dinosaur remains were recovered from it as early as 1915, including partial theropod remains. Weishampel, David B.; ''et al''. (2004). "Dinosaur distributi ...
of the
Buryat Autonomous Soviet Socialist Republic The Buryat Autonomous Soviet Socialist Republic, abbreviated as Buryat ASSR, was an autonomous republic of the Russian SFSR within the Soviet Union. History In May 1923, the republic was created with the name Buryat-Mongol Autonomous Sovie ...
,
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
, dating to the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
period (
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
to
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
stages). It is poorly described, so its relationships cannot be accurately determined (Chure, 2000) and its placement as a species of ''Chilantaisaurus'' is highly questionable. "Chilantaisaurus" ''maortuensis'' was reclassified as ''Shaochilong maortuensis'' in 2009.Brusatte, S., Benson, R., Chure, D., Xu, X., Sullivan, C., and Hone, D. (2009). "The first definitive carcharodontosaurid (Dinosauria: Theropoda) from Asia and the delayed ascent of tyrannosaurids." ''Naturwissenschaften'', An additional species named in 1979, "Chilantaisaurus" ''zheziangensis'', based on bones from the foot and a partial tibia, is actually a
therizinosaur Therizinosaurs (; once called segnosaurs) are an extinct group of large herbivorous theropod dinosaurs whose fossils have been mainly discovered from Cretaceous deposits in Asia and North America. Potential fragmentary remains have also been foun ...
taxon. The cladogram below follows a 2016 analysis by Sebastián Apesteguía, Nathan D. Smith, Rubén Juarez Valieri, and Peter J. Makovicky based on the dataset of Carrano et al. (2012).


References


External links

* {{Taxonbar, from=Q134633 Tetanurae Dinosaur genera Turonian dinosaurs Fossil taxa described in 1964 Dinosaurs of China