Cheilotrema
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''Cheilotrema'' is a genus of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
es belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Sciaenidae Sciaenidae is a family (biology), family of ray-finned fishes belonging to the Order (biology), order Acanthuriformes. They are commonly called drums or croakers in reference to the repetitive throbbing or drumming sounds they make. The family co ...
, the drums and croakers. These fishes are found in the eastern Pacific Ocean.


Taxonomy

''Cheilotrema'' was first proposed as a
monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
in 1861 by the Swiss
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
and
explorer Exploration is the process of exploring, an activity which has some Expectation (epistemic), expectation of Discovery (observation), discovery. Organised exploration is largely a human activity, but exploratory activity is common to most organis ...
Johann Jakob von Tschudi. Tschudi was describing a new species from Peru, ''Cheilotrema fasciatum'', and created the new genus for it. ''Cheilotrema'' belongs to the family Sciaenidae in the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Acanthuriformes Acanthuriformes is a large, diverse order of mostly marine ray-finned fishes, part of the Percomorpha clade. In the past, members of this clade were placed in the suborders Acanthuroidea and Percoidea of the order Perciformes, but this treatment ...
.


Etymology

''Cheilotrema'' combines ''cheilos'', meaning "lip", with ''trema'', which means "pore", an allusion to the obvious pores on the upper lip of ''C. fasciatum''.


Species

''Cheilotrema'' has two recognised species classified within it: * '' Cheilotrema fasciatum'' Tschudi, 1864 (Arnillo drum, Burro) * '' Cheilotrema saturnum'' ( Girard, 1858) (Black croaker)


Characteristics

''Cheilotrema'' croakers have deep, oblong shaped bodies with a sloped head and pointed snout. The mouth is just below the snout and there are 5 pores on the chin but no
barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprini ...
s. The
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
may have either a serrated or smooth margin. The
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
has a deep incision dividing it and there are 10 spines to the front of the incision and a single spine and between 24 and 28 soft rays to its rear. The short anal spine is supported by 2 spines, the second spine being robust and equal in length to the first soft rays, and between 6 and 9 soft rays. The
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
s are short and the
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
is truncate> the body is covered in
ctenoid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...
s apart from around the eyes where there are
cycloid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scale (zoology), scales, which can also provide effective Underwater camouflage, camouflage through the us ...
s. The inner third of the soft rayed part of the dorsal fin and the inner three quarters of the anal fin are scaled. The largest species is the black croaker (''C. saturnum'') which has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of while that of the Arnillo drum (''C. fasciatum'') is .


Distribution and habitat

''Cheilotrema'' croakers are found in the eastern Pacific Ocean, the black croaker is found off western North America off
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
south to southern
Baja California Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...
and in the northern
Gulf of California The Gulf of California (), also known as the Sea of Cortés (''Mar de Cortés'') or Sea of Cortez, or less commonly as the Vermilion Sea (''Mar Vermejo''), is a marginal sea of the Pacific Ocean that separates the Baja California peninsula from ...
. The arnillo drum is found off the western coast of South America from northern Peru to Chile, and around the Galapagos Islands of Ecuador. The black croaker is a demersal fish that is frequently observed in caves and crevices of exposed coasts and open bays in California down to depths of . The allopatric population which inhabits the northern Gulf of California is found over sandy/muddy bottoms. The arnillo drum is described as a sublittoral, demersal fish inhabiting cool waters.


Conservation status

''Cheilotrema'' croakers are classified by the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
, the black croaker is classified as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
, although the population in the Gulf of California is threatened by being taken is
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
in the shrimp trawl and gillnet fisheries there, and there is no information about fisheries for the arnillo drum so it is classified as
Data Deficient A data deficient (DD) species is one which has been categorized by the International Union for Conservation of Nature (IUCN) as offering insufficient information for a proper assessment of conservation status to be made. This does not necessaril ...
.


References

{{taxonbar, from=Q2962149 Sciaenidae Taxa named by Johann Jakob von Tschudi