''Charaxes lycurgus'', the Laodice untailed charaxes, is a

butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...

in the family

Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species ha ...

. It is found in

Guinea Guinea, officially the Republic of Guinea, is a coastal country in West Africa. It borders the Atlantic Ocean to the west, Guinea-Bissau to the northwest, Senegal to the north, Mali to the northeast, Côte d'Ivoire to the southeast, and Sier ...

Sierra Leone Sierra Leone, officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered to the southeast by Liberia and by Guinea to the north. Sierra Leone's land area is . It has a tropical climate and envi ...

Liberia Liberia, officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coast–Lib ...

Ivory Coast Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital city of Yamoussoukro is located in the centre of the country, while its largest List of ci ...

Ghana Ghana, officially the Republic of Ghana, is a country in West Africa. It is situated along the Gulf of Guinea and the Atlantic Ocean to the south, and shares borders with Côte d’Ivoire to the west, Burkina Faso to the north, and Togo to t ...

Togo Togo, officially the Togolese Republic, is a country in West Africa. It is bordered by Ghana to Ghana–Togo border, the west, Benin to Benin–Togo border, the east and Burkina Faso to Burkina Faso–Togo border, the north. It is one of the le ...

Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...

Cameroon Cameroon, officially the Republic of Cameroon, is a country in Central Africa. It shares boundaries with Nigeria to the west and north, Chad to the northeast, the Central African Republic to the east, and Equatorial Guinea, Gabon, and the R ...

Equatorial Guinea Equatorial Guinea, officially the Republic of Equatorial Guinea, is a country on the west coast of Central Africa. It has an area of . Formerly the colony of Spanish Guinea, its post-independence name refers to its location both near the Equ ...

Gabon Gabon ( ; ), officially the Gabonese Republic (), is a country on the Atlantic coast of Central Africa, on the equator, bordered by Equatorial Guinea to the northwest, Cameroon to the north, the Republic of the Congo to the east and south, and ...

, the

Republic of the Congo The Republic of the Congo, also known as Congo-Brazzaville, the Congo Republic or simply the Congo (the last ambiguously also referring to the neighbouring Democratic Republic of the Congo), is a country located on the western coast of Central ...

, the

Central African Republic The Central African Republic (CAR) is a landlocked country in Central Africa. It is bordered by Chad to Central African Republic–Chad border, the north, Sudan to Central African Republic–Sudan border, the northeast, South Sudan to Central ...

Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...

, the

Democratic Republic of the Congo The Democratic Republic of the Congo (DRC), also known as the DR Congo, Congo-Kinshasa, or simply the Congo (the last ambiguously also referring to the neighbouring Republic of the Congo), is a country in Central Africa. By land area, it is t ...

Sudan Sudan, officially the Republic of the Sudan, is a country in Northeast Africa. It borders the Central African Republic to the southwest, Chad to the west, Libya to the northwest, Egypt to the north, the Red Sea to the east, Eritrea and Ethiopi ...

Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...

and

Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...

. The habitat consists of lowland evergreen forests and riverine forests. The larvae feed on '' Pterocarpus santalinoides'', '' Paullinia pinnata'', '' Albizia zygia'', '' Lonchocarpus cyanescens'', ''

Dalbergia ''Dalbergia'' is a large genus of small to medium-size trees, shrubs and lianas in the pea family, Fabaceae, subfamily Faboideae. It was recently assigned to the informal monophyletic ''Dalbergia'' clade (or tribe): the Dalbergieae. The ...

'', ''

Millettia ''Millettia'' is a genus of flowering plants in the family Fabaceae. It consists of about 169 species of shrubs, lianas or trees, which are native to tropical and subtropical regions of sub-Saharan Africa, the Indian subcontinent, Indochina, sout ...

'', ''

Dichapetalum ''Dichapetalum'' is a genus in the plant family Dichapetalaceae. The plants are tropical lianas native mainly to tropical regions of Africa, Asia, Malesia, the West Indies, Australia and Latin America. Some species are known to be poisonous due ...

'' and '' Trachyphrynium'' species.

Subspecies

* ''Charaxes lycurgus lycurgus'' (Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, western Nigeria) * ''Charaxes lycurgus bernardiana'' Plantrou, 1978 (northern and eastern Nigeria, Cameroon, Gabon, Bioko, Congo, Central African Republic, northern Angola, Democratic Republic of the Congo, southern Sudan, western Uganda, Tanzania: eastern shores of Lake Tanganyika)

Taxonomy

Differs from ''

Charaxes zelica ''Charaxes zelica'', the zelica untailed charaxes, is a butterfly in the family Nymphalidae. It is found in Guinea, Ivory Coast, Ghana, Nigeria, Cameroon, Gabon, the Republic of the Congo, the Central African Republic, the Democratic Republic of ...

'' in the deeply concave margin of the forewing, the hindwing blue spots are closer to the margin and the presence of submarginal blue spots on the forewing Kielland, J. 1990 ''Butterflies of Tanzania''. Hill House, Melbourne and London: 1-363. ''Charaxes lycurgus '' is a member of the

species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

''Charaxes lycurgus''. The supposed

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

members are: Clade 1 *''Charaxes lycurgus'' nominate *''

Charaxes porthos ''Charaxes porthos'', the Porthos untailed charaxes, is a butterfly in the family Nymphalidae. It is found in Guinea, Sierra Leone, Ivory Coast, Ghana, Nigeria, Cameroon, the Central African Republic, Gabon, the Republic of the Congo, the Democra ...

'' *''

'' Clade 2 *''

Charaxes mycerina ''Charaxes mycerina'', the mycerina untailed charaxes, is a butterfly in the family Nymphalidae. It is found in Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon, the Central African Republic, the Republic of the Congo, Gabon, the Democ ...

'' *''

Charaxes doubledayi ''Charaxes doubledayi'', Doubleday's untailed charaxes, is a butterfly in the family Nymphalidae. It is found in Sierra Leone, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Bioko, the Republic of the Congo, the Central African Republic, the Democr ...

References

Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentis ...

, 1974 Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part IX. ''Bulletin of the British Museum of Natural History'' (Entomology) 29 (8):415-487

as ''laodice''

External links

Images of ''C. lycurgus lycurgus''

Royal Museum for Central Africa The Royal Museum for Central Africa (RMCA) (; ; ), communicating under the name AfricaMuseum since 2018, is an ethnography and natural history museum situated in Tervuren in Flemish Brabant, Belgium, just outside Brussels. It was originally b ...

(Albertine Rift Project)
''Charaxes lycurgus'' images
at

Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...

''Charaxes lycurgus lycurgus'' images
at BOLD
''Charaxes lycurgus bernardiana'' images
at BOLD Butterflies described in 1793

lycurgus Lycurgus (; ) was the legendary lawgiver of Sparta, credited with the formation of its (), involving political, economic, and social reforms to produce a military-oriented Spartan society in accordance with the Delphic oracle. The Spartans i ...

Butterflies of Africa Taxa named by Johan Christian Fabricius {{Charaxinae-stub