Charassognathus Gracilis
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''Charassognathus'' (meaning 'notched jaw') is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...
cynodont Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 Megaannum, mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extin ...
s. Described in 2007 from a locality near
Fraserburg Fraserburg is a town in the Karoo region of South Africa's Northern Cape province. It is located in the Karoo Hoogland Local Municipality. The town has some of the coldest winters in South Africa. The nearest towns are Williston, Northern Cape, W ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, ''Charassognathus'' is one of the earliest and most basal cynodonts. It is known only from the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, which dates from the
Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...
Period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
. The
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
and only species is ''C. gracilis''. The holotype ( SAM-PK-K 10369), found in the ''Tropidostoma'' Assemblage Zone of the Teekloof Formation, is made up of a crushed
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
, partial lower
jaw The jaws are a pair of opposable articulated structures at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth ...
and one leg. ''Charassognathus'' was a
quadrupedal Quadrupedalism is a form of Animal locomotion, locomotion in which animals have four legs that are used to weight-bearing, bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four l ...
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
. It was named for a notch on its coronoid process which most likely was the insertion point for a chewing muscle, the adductor mandibulae externus. ''Charassognathus'' was a small animal, with a skull only in length. Since the body of ''Charassognathus'' has not been discovered, its full length remains unknown, but estimates have been made at .


Description

''Charassognathus'' has a snout that makes up slightly less than half of the total length of its skull and a long facial process on its septomaxilla. Other than these two features its skull is that of a typical cynodont. The odd shape of its septomaxilla is more typical of
therocephalian Therocephalia is an extinct clade of therapsids (mammals and their close extinct relatives) from the Permian and Triassic periods. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their te ...
s than other
cynodonts Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ances ...
indicating that it may be close to a common ancestor between the two groups.


See also

*
List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all Genus, genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also gene ...


References

* Charassognathidae Lopingian life Lopingian synapsids of Africa Permian South Africa Fossils of South Africa Fossil taxa described in 2007 {{paleo-cynodont-stub