''Changchengornis'' is an extinct basal

pygostylian Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces. Definition The group Pygostylia was intended to encompass all avialans with a short, stubby tail, as opposed to t ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

from the Early

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

. Its remains have been found in the

People's Republic of China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, sli ...

, in

Chaomidianzi Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...

rocks from around the

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma ( million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is pre ...

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...

boundary, deposited 125 million years ago. ''Changchengornis'' was a close relative of the better-known ''Confuciusornis''. In 1999 it was assigned to the

Confuciusornithidae Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their cl ...

. The more pointed bill of ''Changchengornis'' might indicate a diet different from that of ''Confuciusornis''. However, of ''Confuciusornis'' itself it is contested whether it were a fish eater, an

omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...

, or a seed eater.

Discovery

In 1998

Ji Shuan Ji or JI may refer to: Names and titles * Ji (surname), the pinyin romanization of a number of distinct Chinese surnames * Ji (Korean name), a Korean surname and element in given names (including lists of people with the name) * -ji, an honorif ...

and

Luis Chiappe Luis María Chiappe (born 18 June 1962) is an Argentine paleontologist born in Buenos Aires who is best known for his discovery of the first sauropod nesting sites in the badlands of Patagonia in 1997 and for his work on the origin and early evol ...

discovered among the many specimens of the National Geological Museum of China at

Beijing } Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 21 ...

assigned to ''

Confuciusornis ''Confuciusornis'' is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, ''Confuciusornis'' had a toothless beak, b ...

'', an exemplar that seemed somewhat different. Subsequent preparation by the American Museum of Natural History showed that it indeed was a separate species, new to science. In 1999 Ji, Chiappe and

Ji Qiang Ji or JI may refer to: Names and titles * Ji (surname), the pinyin romanization of a number of distinct Chinese surnames * Ji (Korean name), a Korean surname and element in given names (including lists of people with the name) * -ji, an honorific ...

named the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

and only species of ''Changchengornis'': ''Changchengornis hengdaoziensis''. The generic name refers to the

Great Wall of China The Great Wall of China (, literally "ten thousand ''li'' wall") is a series of fortifications that were built across the historical northern borders of ancient Chinese states and Imperial China as protection against various nomadic grou ...

, ''changcheng'', and combines this with a Greek ὄρνις, ''ornis'', "bird". The specific name refers to the geological Hengdaozi Member. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

specimen, GMV-2129, was found near the village of Jianshangou in Liaoning province, in the Jianshangou Beds of the Hengdaozi Member of the

Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ...

, at the time seen as a separate Chaomidianzi Formation. It consists of a plate and counterplate, GMV-2129a/b, showing a largely complete and articulated but compressed and somewhat damaged skeleton. Much of the feathering has been preserved in an excellent state of preservation.

Description

''Changchengornis'' resembles its relative ''Confuciusornis''. The type specimen is rather small, smaller than the smallest known specimens of ''Confuciusornis''. Compared to the latter, ''Changchengornis'' had a beak that was more pointed, slightly hooked at the tip, proportionally shorter, and higher at the back.Mortimer, Michael (2004): The Theropod Database
Phylogeny of taxa
. Retrieved 2013-MAR-02.) The deltopectoral crest of the

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

is not pierced. GMV-2129 also shows two elongated, ribbon-like tail feathers as found in some specimens of ''Confuciusornis'', that are often considered to be the males. Also, the fossil gives the impression of a head tuft or crest being present; if so, the outline of its head must have borne an uncanny resemblance to today's ''

Tauraco ''Tauraco'' is a genus of turacos. It contains the "typical" or green turacos; though their plumage is not always green all over, the presence of significant amounts of turacoverdin-colored plumage generally sets ''Tauraco'' species apart from ot ...

'' or

turaco The turacos make up the bird family Musophagidae ( "banana-eaters"), which includes plantain-eaters and go-away-birds. In southern Africa both turacos and go-away-birds are commonly known as loeries. They are semi-zygodactylous: the fourth ( ...

s in general but it could also be an artefact of conservation.

References

{{Taxonbar, from=Q2425376 Confuciusornithids Early Cretaceous birds of Asia Fossil taxa described in 1999 Yixian fauna