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The Chacoan peccary or ''tagua'' (''Catagonus wagneri'' or ''Parachoerus wagneri'') is the last extant species of the genus ''Catagonus''; it is a peccary found in the Gran Chaco of
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
, and
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. Approximately 3,000 remain in the world.


History

The Chacoan peccary was first described in 1930 based on
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s and was originally thought to be an extinct species. In 1971, the animal was discovered to still be alive in the Chaco region, in the Argentine province of
Salta Salta () is the capital and largest city in the Provinces of Argentina, Argentine province of Salta Province, the same name. With a population of 618,375 according to the 2010 census, it is also the List of cities in Argentina, 7th most-populous ...
. The species was well-known to the native people, but it took a while for Western scientists to acknowledge its existence; it is known locally as the ''tagua''. Because it was originally described as extinct before its "rediscovery", it is an example of a Lazarus taxon.


Taxonomy

The Chacoan peccary is notable in that it is not the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of its genus, ''Catagonus'', despite being the only living representative. Instead, the type is the extinct '' Catagonus metropolitanus''. Such a case is an example of a Lazarus taxon, and shares this trait with another South American native, the bush dog. The Chacoan peccary was first described by scientists in 1972. A 2017 study on the phylogenetic systematics of Tayassuidae species suggests that ''Catagonus'' should only contain ''C. metropolitanus''. The extinct narrow-headed peccary ''(C. stenocephalus)'' should be moved into ''Brasiliochoerus'', while the Chacoan peccary, ''Catagonus bonaerensis'' and '' Catagonus carlesi'' should be placed in ''Parachoerus''. If this is accepted, then ''Catagonus'' becomes an extinct genus once more.


Habitat

The Chacoan peccary is confined to hot, dry areas. Dominated by low-lying succulents and thorny bushes, the Gran Chaco is about 140,000 km2. A few scattered giant trees are found, but the majority of the vegetation is thorny scrub vegetation. The Chacoan peccary has developed adaptations such as well-developed sinuses to combat dry, dusty conditions. Their feet are also small, which allows maneuverability among spiny plants.


Physical characteristics

The largest of the three generally accepted species of peccaries, the Chacoan peccary has many pig-like features. It is an
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
with a well-formed rostrum with a tough leathery snout. The bristle-like hair is generally brown to almost gray. A dark stripe runs across the back, and white fur is on the shoulders. Chacoan peccaries differ from other peccary species by having longer ears, snouts, and tails. It has white hairs around the mouth, unlike other peccaries. ''Catagonus wagneri'' also has a third hind toe, but other peccaries only have two. The hypsodont teeth follow this dental formula: 2/3, 1/1, 3/3, 3/3. The upper canines display the distinguishing trait of peccaries, pointing downwards instead of out and up like pigs, which may be a signal for other peccaries to keep the group together through the dense bush.Catagonus wagneri - Chacoan peccary


Reproduction

Young are generally born between September and December, but litters have been found almost year-round. Births have been linked to periods of food abundance and rainfall. The average number of embryos has been recorded as approximately 2.72. Females may leave the herd to give birth and then return afterwards. Newborns are
precocial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
, able to run a few hours after birth. The pelage of the young resembles that of the adults. There is no
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
.


Behavior

Chacoan peccaries often travel in herds of up to 20 individuals. They are active during the day, especially in the morning when they are most apt to travel. Herds display a general travel cycle within their home range of 42 days. This allows the individuals to monitor and show ownership over their areas. These social mammals communicate by various sounds, ranging from grunts to chatters of the teeth. Though individuals may occasionally exhibit aggressive behavior such as charging and biting, this species is not as aggressive as others. As a defensive strategy, members of a herd may line up in a defensive wall; this makes the herds harder targets for hunters. The Chacoan peccary produces a milky, odorous substance used for marking trees, shrubs, and similar. The substance is secreted from glands located on their backs, and is dispersed by rubbing. Frequently bathing in mud or dust, Chacoan peccaries also defecate at particular "stations".


Food habits

The arid habitat of the Gran Chaco region provides very tough vegetation for the Chacoan peccary. These peccaries feed on various species of cacti, such as '' Cleistocactus baumannii'' and '' Opuntia discolor''. It uses its tough snout to roll the cacti on the ground, rubbing the spines off. It may pull off the spines with its teeth and spit them out. The kidneys are specialized to break down acids from the cacti. The two-chambered stomachs are also well suited to digest tough foods. Occasionally grazing on
bromeliad The Bromeliaceae (the bromeliads) are a Family (biology), family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the Tropics, tropical Americas, with several species found in the American subtropics and on ...
roots, it also eats acacia pods and fallen cactus flowers. This species of peccary seeks out salt licks formed from ant mounds and construction projects (road building and land clearings). The Chacoan peccary gains essential minerals like calcium,
magnesium Magnesium is a chemical element; it has Symbol (chemistry), symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 ...
, and
chlorine Chlorine is a chemical element; it has Symbol (chemistry), symbol Cl and atomic number 17. The second-lightest of the halogens, it appears between fluorine and bromine in the periodic table and its properties are mostly intermediate between ...
from the salt licks.


Conservation status

Because the Chacoan peccary is endemic to a formerly isolated region of South America, it is most vulnerable to human activity. Just as quickly as this species is discovered in an area, it disappears. Herd numbers are decreasing as a result of
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and fragmentation. Their range is being quickly transformed into large Texas-style
ranch A ranch (from /Mexican Spanish) is an area of landscape, land, including various structures, given primarily to ranching, the practice of raising grazing livestock such as cattle and sheep. It is a subtype of farm. These terms are most often ap ...
es. Hunting also continues, as well as an unidentified disease that has plagued the herds in recent years. A population has been established in North American and European zoos. Preserves have also been established in Paraguay, but are not highly enforced.


References


Further reading

* * * * {{Authority control Peccaries Mammals of Argentina Mammals of Bolivia Mammals of Paraguay Gran Chaco EDGE species Mammals described in 1930