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The Cerrejón Formation is a geologic formation in Colombia dating back to the Middle-Late
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
. It is found in the El Cerrejón sub-basin of the Cesar-Ranchería Basin of La Guajira and
Cesar Cesar, César or Cèsar may refer to: Arts, entertainment, and media * ''César'' (film), a 1936 film directed by Marcel Pagnol * ''César'' (play), a play by Marcel Pagnolt * César Award, a French film award Places * Cesar, Portugal * Ces ...
. The formation consists of
bituminous coal Bituminous coal, or black coal, is a type of coal containing a tar-like substance called bitumen or asphalt. Its coloration can be black or sometimes dark brown; often there are well-defined bands of bright and dull material within the coal seam, ...
fields that are an important economic resource. Coal from the Cerrejón Formation is mined extensively from the
Cerrejón Cerrejón is a large open-pit coal mine in Northern Colombia owned by Glencore. At Cerrejón, low-ash, low-sulphur bituminous coal from the Cerrejón Formation is excavated. At over the mine is one of the largest of its type, the largest in La ...
open-pit Open-pit mining, also known as open-cast or open-cut mining and in larger contexts mega-mining, is a surface mining technique of extracting rock or minerals from the earth from an open-air pit, sometimes known as a borrow. This form of mining ...
coal mine, one of the largest in the world. The formation also bears fossils that are the earliest record of
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfo ...
s.Wing et al., 2009


Definition

The formation was first named Septarias Formation and in 1958 renamed to Cerrejón Formation by
Thomas van der Hammen Thomas van der Hammen (Schiedam, Netherlands, 27 September 1924 - Chía, Colombia, 12 March 2010) was a Dutch palaeontologist, botanist and geologist. He had published more than 160 works in five languages.Los Cuervos LOS, or Los, or LoS may refer to: Science and technology * Length of stay, the duration of a single episode of hospitalisation * Level of service, a measure used by traffic engineers * Level of significance, a measure of statistical significance ...
and
Bogotá Formation The Bogotá Formation ( es, Formación Bogotá, E1-2b, Tpb, Pgb) is a geological formation of the Eastern Hills and Bogotá savanna on the Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The predominantly shale and siltstone for ...
s to the south;
Llanos Orientales The Llanos ( Spanish ''Los Llanos'', "The Plains"; ) is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela, in northwestern South America. It is an ecoregion of the tropical and subtropical grassland ...
and
Altiplano Cundiboyacense The Altiplano Cundiboyacense () is a high plateau located in the Eastern Cordillera of the Colombian Andes covering parts of the departments of Cundinamarca and Boyacá. The altiplano corresponds to the ancient territory of the Muisca. The Alt ...
respectively. The formation is also time-equivalent with the Marcelina Formation of the Venezuelan
Serranía del Perijá The Serranía del Perijá, Cordillera de Perijá or Sierra de Perijá is a mountain range, an extension of the eastern Andean branch ( Cordillera Oriental), in northern South America, between Colombia and Venezuela, ending further north in the ...
and the Catatumbo Formation of the southwestern Colombian part of the
Maracaibo Basin The Maracaibo Basin, also known as Lake Maracaibo natural region, Lake Maracaibo depression or Lake Maracaibo Lowlands, is a foreland basin and one of the eight natural regions of Venezuela, found in the northwestern corner of Venezuela in South A ...
, the Catatumbo Basin.Ojeda & Sánchez, 2013, p.69 The formation has also been described as laterally equivalent to the Mostrencos and Santa Cruz Formations of Venezuela.Rodríguez & Londoño, 2002, p.165 Based on
lithofacies Lithofacies may refer to: * A mappable subdivision of a designated stratigraphic unit, distinguished from adjacent subdivisions on the basis of lithology; a facies characterized by particular lithologic features * The rock record of any particular ...
associations and paleofloral composition, the
depositional environment In geology, depositional environment or sedimentary environment describes the combination of physical, chemical, and biological processes associated with the deposition of a particular type of sediment and, therefore, the rock types that will b ...
fluctuated from an
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environmen ...
-influenced
coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Co ...
at the base of the formation to a
fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluv ...
-influenced coastal plain at the top. In the geologically recent past, some coal in the formation has spontaneously and naturally combusted to form clinker, red and brick-looking burnt coal. These rocks outcrop irregularly and are up to thick. Clinker is found near deformed zones such as faults or tight folds, and is older than the deformities themselves. They are thought to have combusted after the development of the Cerrejón
thrust fault A thrust fault is a break in the Earth's crust, across which older rocks are pushed above younger rocks. Thrust geometry and nomenclature Reverse faults A thrust fault is a type of reverse fault that has a dip of 45 degrees or less. If ...
and
alluvial fan An alluvial fan is an accumulation of sediments that fans outwards from a concentrated source of sediments, such as a narrow canyon emerging from an escarpment. They are characteristic of mountainous terrain in arid to semiarid climates, but a ...
.


Paleoenvironment

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s found from the Cerrejón Formation are the earliest record of Neotropical rainforests, with an abundance of plant macrofossils and palynomorphs. The Cerrejón Formation also records a riverine vertebrate fauna that includes
lungfish Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, i ...
, turtles, snakes, and crocodyliforms. Based on these fossils and the stratigraphy of the formation, the Cerrejón Formation was likely formed on a coastal plain, covered in a wet tropical rainforest and incised by a large river system.Head et al., 2009 The rainforest is estimated to have been around 5°N paleolatitude. During the Paleocene, equatorial temperatures were much higher than they are today. Based on the size of the giant boid '' Titanoboa'', specimens of which have been found in the Cerrejón Formation, the mean annual temperature of Paleocene equatorial South America was between and . This is the minimum annual temperature range that a
poikilotherm A poikilotherm () is an animal whose internal temperature varies considerably. Poikilotherms have to survive and adapt to environmental stress. One of the most important stressors is temperature change, which can lead to alterations in membrane ...
as large as ''Titanoboa'' could live. It is consistent with Paleocene climate models which predict greenhouse temperatures and an atmospheric pCO2 concentration of around 2,000 parts per million. Paleotemperature estimates based on fossil leaf assemblages from the Cerrejón Formation predict the mean annual temperature to be lower than other estimates. However, such temperature estimates based on
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
and
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
rainforest paleoflora have been considered underestimates. Mean annual temperatures of are considered to be too high for modern tropical forests, but the Cerrejón rainforest could have been maintained by increased atmospheric pCO2 levels and the high regional rainfall, which is estimated to have been around per year.


Flora

The floral record of the Cerrejón Formation is well known, with many identifiable and well preserved plant microfossils having been found from the Cerrejón mine. The fossils are well preserved, and in some cases their cell structure is intact. In comparison to modern Neotropical rainforests, the diversity of plants is quite low. This may be an indication of the early stage of Neotropical diversification, or a delayed recovery period following the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
. Many plants from the Cerrejón Formation belong to families that are still common today in modern Neotropical rainforests. There is a diverse variety of palms and
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
s in the formation. In addition to palms and legumes, much of the biomass of the Paleocene forest consisted of
laurales The Laurales are an order of flowering plants. They are magnoliids, related to the Magnoliales. The order includes about 2500-2800 species from 85-90 genera, which comprise seven families of trees and shrubs. Most of the species are tropical a ...
,
malvales The Malvales are an order of flowering plants. As circumscribed by APG II-system, the order includes about 6000 species within 9 families. The order is placed in the eurosids II, which are part of the eudicots. The plants are mostly shrubs a ...
, menisperms, aroids, and zingiberaleans. Studies of fossil plants from
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
-age sites indicate that the floral composition below the
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end ...
(K–T boundary) was very different from that of the Paleocene. Legumes are absent from Cretaceous strata, and likely appeared or diversified during the Paleocene. The presence of these types of flora in Paleocene strata shows that plants characteristic of modern Neotropical rainforests have existed for geologically long periods of time, being able to withstand climatic and geographic changes in South America. It has been suggested that today's Neotropical rainforests are the result of environmental changes brought about by Quaternary glacial cycles (i.e. the recent ice age). These cycles would have caused fluctuation in the diversity and extent of rainforests. If this was the case, the current diversity of the Amazon rainforest would be a recent speciation in a changing environment. However, the floral record from the Cerrejón Formation shows that the current diversity of the Amazon Rainforest can be traced back into the early Cenozoic.


Fauna

Feeding damage from insects is evident on some of the plant macrofossils from the Cerrejón Formation. One survey of plant macrofossils showed that around half of the studied specimens had been attacked by herbivorous insects. The insects that damaged the leaves were predominantly generalist feeders, unlike modern Neotropical insects that are mainly
specialist Specialist may refer to: Occupations * Specialist (rank), a military rank ** Specialist (Singapore) * Specialist (arena football) * Specialist degree, in academia * Specialty (medicine) * Designated market maker, in the American stock market * ...
herbivores. There is no evidence of the elevated insect-feeding diversity or host-specialized feeding associations that are seen in later
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...
forests. The insect diversity in the Cerrejón Formation is low in comparison to the diversity of insects in Neotropical rainforests today, and it is likely that leaf damage was made by relatively few species. Remains of the giant boid '' Titanoboa cerrejonensis'' have been found from a gray
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...
layer underlying Coal Seam 90 in the Cerrejón mine. ''Titanoboa'' is the largest known snake to have ever existed, reaching an estimated length of . ''Eunectes'', the anaconda, is likely to be a close living analogue of ''Titanoboa''. A dyrosaurid
crocodylomorph Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cro ...
called ''
Cerrejonisuchus improcerus ''Cerrejonisuchus'' is an extinct genus of dyrosaurid crocodylomorph. It is known from a complete skull and mandible from the Cerrejón Formation in northeastern Colombia, which is Paleocene in age. Specimens belonging to ''Cerrejonisuchus'' and ...
'' was described in 2010 from the Cerrejón Formation in the same layer as ''Titanoboa''. It was a small dyrosaurid, and had the shortest snout length relative to its skull length of any dyrosaurid. Most dyrosaurids were marine, with long snouts adapted for catching fish. The short snout of ''Cerrejonisuchus'' is likely to have been an adaptation for a more generalized diet in a transitional aquatic environment. It is possible that ''Cerrejonisuchus'' was a food source for ''Titanoboa'', as the two inhabited the same riverine environment. The anaconda has been documented consuming
caiman A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South Ameri ...
s, a feeding habit that is similar to the inferred habit of ''Titanoboa''. A second dyrosaurid, '' Acherontisuchus'', was named in 2011 from the formation. With a large body and long snout, it resembles most other dyrosaurids.Hastings et al., 2011 A third dyrosaurid, '' Anthracosuchus'', was named in 2014. Unlike other dyrosaurids, it had blunt teeth and a short skull. ''Anthracosuchus'' likely had massive jaw muscles allowing it to feed on large turtles, a behavior which is corroborated by predation marks found at the site.


Coal resources

The Cerrejón Formation contains extensive coal seams that are mined primarily at the Cerrejón mine. The coal is desirable for its low
ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...
and
sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formul ...
content and for its resistance to caking. Cerrejón is Colombia's largest coal producing mine, with most of the production exported to Europe. It is the largest coal mining operation in Latin America, with an estimated 28.4 million tons mined in 2006.


Itaboraian correlations


References


Bibliography

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Further reading

* * * * * {{DEFAULTSORT:Cerrejon Formation Geologic formations of Colombia Formations Paleogene Colombia Paleocene Series of South America Itaboraian Peligran Thanetian Stage Coal in Colombia Coal formations Mudstone formations Sandstone formations Siltstone formations Fluvial deposits Reservoir rock formations Source rock formations Formations Formations