Ceratophyllaceae
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Ceratophyllaceae is a
cosmopolitan Cosmopolitan may refer to: Internationalism * World citizen, one who eschews traditional geopolitical divisions derived from national citizenship * Cosmopolitanism, the idea that all of humanity belongs to a single moral community * Cosmopolitan ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s including one living genus commonly found in ponds, marshes, and quiet streams in tropical and in temperate regions. It is the only
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
family in the order Ceratophyllales. Species are commonly called coontails or hornworts, although hornwort is also used for unrelated plants of the division
Anthocerotophyta Hornworts are a group of non-vascular Embryophytes (land plants) constituting the division Anthocerotophyta (). The common name refers to the elongated horn-like structure, which is the sporophyte. As in mosses and liverworts, hornworts have a ...
. Living ''Ceratophyllum'' grows completely submerged, usually, though not always, floating on the surface, and does not tolerate drought.


Taxonomy

Ceratophyllaceae was considered a relative of
Nymphaeaceae Nymphaeaceae () is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate climate, temperate and tropical climates around the world. The family contains five genera with about 70 know ...
and included in
Nymphaeales The Nymphaeales are an order of flowering plants, consisting of three families of aquatic plants, the Hydatellaceae, the Cabombaceae, and the Nymphaeaceae (water lilies). It is one of the three orders of basal angiosperms, an early-divergin ...
in the
Cronquist system The Cronquist system is a list of systems of plant taxonomy, taxonomic classification system of angiosperms, flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of ...
, but research has shown that it is not closely related to Nymphaeaceae or any other extant plant family. Some early molecular phylogenies suggested it was the
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to all other angiosperms, but more recent research suggests that it is the sister group to the
eudicots The eudicots or eudicotyledons are flowering plants that have two seed leaves (cotyledons) upon germination. The term derives from ''dicotyledon'' (etymologically, ''eu'' = true; ''di'' = two; ''cotyledon'' = seed leaf). Historically, authors h ...
. The APG III system placed the family in its own order, the Ceratophyllales.Kew World Checklist of Selected Plant Families
/ref>Angiosperm Phylogeny Web

/ref> The
APG IV system The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publish ...
accepts the phylogeny shown below: The extinct family Montsechiaceae containing the genus '' Montsechia'' has also been placed in the order Ceratophyllales.


Genera

The family contains one living genus, and several extinct genera described from the fossil record, including one of the earliest fruit bearing (in the form of an achene) plants, the Dakota formation freshwater genus '' Donlesia'' from Early Cretaceous. *''
Ceratophyllum ''Ceratophyllum'' is a cosmopolitan distribution, cosmopolitan genus of flowering plants commonly found in ponds, marshes, and quiet streams in tropical and in temperate regions. It is the only extant taxon, extant genus in the family Ceratophyll ...
'' *†'' Ceratostratiotes'' *†'' Donlesia''


References

{{Taxonbar, from=Q12012524 Angiosperms Freshwater plants Angiosperm families