''Ceratophaga'' is a genus of

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

s belonging to the family

Tineidae Tineidae is a family of moths in the order Lepidoptera described by Pierre André Latreille in 1810. Collectively, they are known as fungus moths or tineid moths. The family contains considerably more than 3,000 species in more than 300 genera. ...

. The name ''"ceratophaga"'' is derived from the Greek for "horn eater". Sixteen species are currently recognised, widespread in the

Afrotropical realm The Afrotropical realm is one of the Earth's eight biogeographic realms. It includes Sub-Saharan Africa, the southern Arabian Peninsula, the island of Madagascar, and the islands of the western Indian Ocean. It was formerly known as the Ethiopi ...

and

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

. In the Americas one species has been described: '' Ceratophaga vicinella'', which occurs in the southeastern

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

. Twelve of the known species occur in

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

, and of those '' Ceratophaga vastella'' is perhaps the best-known. The

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e of most

(of which clothes moths are the most familiar) have adapted to feeding on non-herbaceous material, but ''Ceratophaga'' are remarkable in that they feed, apparently exclusively, on solid

keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. It is the key structural material making up Scale (anatomy), scales, hair, Nail (anatomy), nails, feathers, horn (anatomy), horns, claws, Hoof, hoove ...

from dead

vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...

s. For most species this usually means the

horns Horns or The Horns may refer to: * Plural of Horn (anatomy) * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * Horns (novel), ''Horns'' (novel), a dar ...

and

hooves The hoof (: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits; the ruminants with ...

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

s, but the North American ''Ceratophaga vicinella'' feeds on the shells of the

tortoise Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...

Gopherus polyphemus The gopher tortoise (''Gopherus polyphemus'') is a species of tortoise in the Family (biology), family Testudinidae. The species is native to the southeastern United States. The gopher tortoise is seen as a keystone species because it digs burrow ...

''.

References

A Caterpillar that Eats Tortoise Shells
Tineidae Tineidae genera {{Tineidae-stub