''Krascheninnikovia'' is a genus of flowering plants in the subfamily

Chenopodioideae The Chenopodioideae are a subfamily of the flowering plant family Amaranthaceae in the APG III system, which is largely based on molecular phylogeny, but were included – together with other subfamilies – in the family Chenopodiaceae, or goose ...

of the family

Amaranthaceae Amaranthaceae ( ) is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, maki ...

known as winterfat, so-called because it is a nutritious livestock forage. They are known from Eurasia and western North America. These are hairy perennials or small

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

s which may be

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...

or dioecious. They bear spike

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

s of woolly flowers.

Description

The species of ''Krascheninnikovia'' are erect

subshrub A subshrub (Latin ''suffrutex'') or undershrub is either a small shrub (e.g. prostrate shrubs) or a perennial that is largely herbaceous but slightly woody at the base (e.g. garden pink and florist's chrysanthemum). The term is often interch ...

s or

shrubs A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

. The plants are densely covered with dendroid stellate hairs and additionally with simple, unbranched hairs. The alternate

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

stand solitary or grouped in fascicles, and can be petiolate or nearly sessile. The flat, non-fleshy leaf blades are linear to narrowly lanceolate to ovate, with entire margins, and truncate, cuneate, rounded, or subcordate base. The flowers are

unisexual Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

, the plants can be

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

. Male flowers form an interrupted spike or subcapitate

of glomeruled, ebracteate flowers. These consist of 4 basally connate

perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...

segments, that are ovate or elliptic, membranous and abaxially hairy; and 4

stamens The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...

with oblong

anther The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s and linear, exserted filaments. Female flowers sitting single or paired axillary, enclosed by 2 hairy

bracteole In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also look ...

s, that are connate in the lower part, compressed to slightly keeled, with 4 hornlike tips; a perianth is missing, the female flowers consist just of an ovary with a short style and 2 elongated stigmas. The hairy fruit enclosed by the bracteoles is ovate and compressed, its membranous pericarp is free from the seed. The vertically orientated seed has a brown thin seed coat covered with white hairs. The embryo is nearly annular or horseshoe-shaped and encloses the copious perisperm. The

chromosome A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most import ...

base number is x=9, for example 2n=36 for ''Krascheninnikovia ceratoides'', 2n=18 and 2n=36 for ''Krascheninnikovia lanata''.

Distribution

The species of ''Krascheninnikovia'' are mostly distributed in

Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...

, two species occur in

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. In

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

, ''

Krascheninnikovia ceratoides ''Krascheninnikovia'' is a genus of flowering plants in the subfamily Chenopodioideae of the family Amaranthaceae known as winterfat, so-called because it is a nutritious livestock forage. They are known from Eurasia and western North America. Th ...

'' is the only species, it is native in eastern and Southern Europe, Winterfat occurs in dry valley bottoms, on flat mesas, and on hillsides, at elevations between 2,400 and 9,300 feet. It is drought resistant and intolerant of flooding, excess water, or acidic soils. Seed production especially in desert regions, is dependent on precipitation. Good seed years occur when there is appreciable summer precipitation and little browsing. Winterfat occurs on well-drained, calcareous soils with low to moderate salt concentrations.

Systematics

''Krascheninnikovia'' has been first described in 1772 by

Johann Anton Güldenstädt Johann Anton Güldenstädt (26 April 1745 in Riga, Latvia – 23 March 1781 in St. Petersburg, Russia) was a Baltic German natural history, naturalist and explorer in Russian service. Güldenstädt lost both his parents early, and from 1763 on ...

. The genus was named for the

Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...

botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

and explorer of

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

and

Kamchatka The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...

Stepan Krasheninnikov Stepan Petrovich Krasheninnikov (; – ) was a Russian explorer of Siberia, naturalist and geographer who gave the first full description of Kamchatka in the early 18th century. He was elected to the Russian Academy of Sciences in 1745. The Kra ...

. The type species is ''

'' (L.) Gueldenst. The older name ''Ceratoides'' had to be rejected: it referred just to a pre-

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

description by

Joseph Pitton de Tournefort Joseph Pitton de Tournefort (5 June 165628 December 1708) was a French botanist, notable as the first to make a clear definition of the concept of genus for plants. Botanist Charles Plumier was his pupil and accompanied him on his voyages. Li ...

, comprising the type of the related genus '' Ceratocarpus''. The genus ''Krascheninnikovia'' belongs to the subfamily

of the family

. After

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

research, it was grouped into the tribe Axyrideae. Krascheninnikovia ceratoides sl 4

The genus includes three accepted species:) *''

'' (including ''Krascheninnikovia lenensis'' ) – from Europe to East Asia * '' Krascheninnikovia fruticulosa'' – in Asia *''

Krascheninnikovia lanata ''Krascheninnikovia lanata'' is a species of flowering plant currently placed in the family Amaranthaceae (previously, Chenopodiaceae), known by the common names winterfat, white sage, and wintersage. It is native to much of western North Americ ...

'' – in North America

References

Pertti Uotila, 2011: ''Chenopodiaceae (pro parte majore):'
''Krascheninnikovia'' – In: ''Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity''
accessed 2013-07-15

External links

Jepson Manual TreatmentUSDA Plants Profile
{{Authority control Chenopodioideae Amaranthaceae genera Taxa named by Johann Anton Güldenstädt Flora of the Holarctic realm