Cephalotaxaceae
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Cephalotaxaceae is a small
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
s. It is often treated as a monophyletic family including only the single genus '' Cephalotaxus'', while others have also included '' Torreya'' and ''
Amentotaxus ''Amentotaxus'' is a genus of Pinophyta, conifers (catkin-yews) comprising five species, treated in either the Cephalotaxaceae, or in the Taxaceae when that family is considered in a broad sense. The genus is endemic (ecology), endemic to subtr ...
'' in the family. Cephalotaxaceae is closely allied to the yew family
Taxaceae Taxaceae (), commonly called the yew family, is a coniferous family (biology), family which includes six Extant taxon, extant and two extinct genera, and about 30 species of plants, or in older interpretations three genera and 7 to 12 species. ...
, and is often included in a wide interpretation of Taxaceae, based on phylogenetic evidence and close morphological similarities between them. Included species were restricted to east
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, except for two species of '' Torreya'' found in the southwest and southeast of the United States;
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
evidence shows a much wider prehistorical
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
distribution. The most notable differences between Taxaceae and Cephalotaxaceae concerned the cone aril, which fully encloses the seeds of Cephalotaxaceae, the longer maturation of Cephalotaxaceae seeds and the larger size of the mature seeds. When considered as a distinct family, members of Cephalotaxaceae are much branched, small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s and
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s. The leaves are
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
, spirally arranged, often twisted at the base to appear biranked. They are linear to lanceolate, and have pale green or white stomatal bands on the undersides. The plants are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...
, subdioecious, or dioecious. The male cones are 4–25 mm long, and shed
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
in the early spring. The female cones are reduced, with one to a few ovuliferous scales, and one seed on each ovuliferous scale. As the seed matures, the ovuliferous scale develops into a fleshy
aril An aril (), also called arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode, or false aril, is sometimes distinguished: whereas an aril grows from the attachment point of the seed to the ova ...
fully enclosing the seed. The mature aril is thin, green, purple or red, soft and resinous. Each ovuliferous scale remains discrete, so the cone develops into a short stem with one to a few
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
-like seeds. They are probably eaten by
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s or other
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s which then disperse the hard seed undamaged in their droppings, but seed dispersal mechanisms in the family are not yet well researched.


References

{{Taxonbar, from=Q841198 Pinales families Historically recognized plant families