Cephalopholis Boenak
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''Cephalopholis boenak'', the chocolate hind, brownbarred rockcod, brown-banded cod or brown-banded rockcod, is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
, a
grouper Groupers are a diverse group of marine ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all members of this f ...
from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Epinephelinae Groupers are a diverse group of marine Actinopterygii, ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all m ...
which is in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...
which also includes the
anthias Anthias are members of the family Anthiadidae in the order Perciformes. The group has also been called Anthiidae or Anthiinae, but these names are preoccupied by a subfamily of ground beetles in the family Carabidae erected by Bonelli in 1813 ...
and sea basses. It is associated with reefs over a wide
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
distribution. It is an important species for commercial fisheries in some parts of its range.


Description

''Cephalopholis boenak'' has a body which has a
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
which is 2.6–3.1 times its depth and with a head that is around a fifth of the standard length. Its
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
contains 9 spines and 15–17 soft rays while the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
has 3 spines and 8 soft rays. The preopercle is rounded with fine serrations but no enlarged spines at its angle and with a flesh lower margin. The body, including the abdomen, is covered in
ctenoid A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...
scales which is a characteristic of this species. The overall colour is brownish to greenish-grey and there are normally 7–8 dark vertical bars on the flanks. The dorsal fin has a wide dark submarginal band and a thin white margin, these are also present on the anal fins, and the upper and lower lobes of the
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
. There is a blackish spot on the upper margin of the gill cover. The juveniles have a yellowish on the posterior portion of their body and are marked with a white stripe which runs from the snout to the origin of the dorsal fin. It grows to a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of .


Distribution

''Cephalopholis boenak'' has a wide distribution in the Indian Ocean and the
Western Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the contine ...
from the eastern coast of Africa where it occurs from Kenya south to southern Mozambique along the southern Asian coast and into the Pacific where it occurs north as far as the
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Geography of Taiwan, Taiwan: the Ryukyu Islands are divided into the Satsunan Islands (Ōsumi Islands, Ōsumi, Tokara Islands, Tokara and A ...
and east to
New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...
. In Australia it has been recorded along the northern coast from
Coral Bay, Western Australia Coral Bay is a small coastal settlement located north of Perth, in the Shire of Carnarvon in the Gascoyne region of Western Australia. Bordered by the Ningaloo Reef, it is a popular tourist destination and largely owes its survival to revenue ...
to
Moreton Bay Moreton Bay is a bay located on the eastern coast of Australia from central Brisbane, Queensland. It is one of Queensland's most important coastal resources. The waters of Moreton Bay are a popular destination for recreational anglers and are ...
in
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
.


Habitat and biology

''Cephalopholis boenak'' is mainly a coastal species which is found on silty, dead and living coral reefs as deep as . It is a carnivore which preys mainly on other fishes and on
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s. The chocolate hind is a
protogynous hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodite ...
, they undergo a sex change from females to males, although some are born as males. They occur in relatively small social groups which comprise a single male, one or two smaller females, and a variable number of sexually inactive individuals. These sexually inactive fish are sexually inactive adults or juveniles. The males defend a
territory A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
from other males and the territory is much larger than the home ranges of the females. The males visit the females and other fish within their territory at specific locations, almost always using the same route. Courtship occurs in the late afternoons but breeding behaviour is not associated with any particular moon phase and can happen multiple times during the breeding season.


Utilisation

''Cephalopholis boenak'' is commercially fished for human consumption in many parts of its range. Hook-and-line, traps and trawling are the main methods used to catch them, although they are occasionally caught using gillnet and purse-seine. In
Hong Kong Hong Kong)., Legally Hong Kong, China in international treaties and organizations. is a special administrative region of China. With 7.5 million residents in a territory, Hong Kong is the fourth most densely populated region in the wor ...
it has increased in commercial importance as a result of the decline of larger-bodied species of grouper and the chocolate hind is now one of the more important species in the Hong Kong fish market. It is commonly landed by artisanal fishermen in southern Kenya. In Sri Lanka it is also harvested to be traded in the marine
aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...
trade.


Taxonomy

''Cephalopholis boenak'' was first formally described as ''Bodianus boenak'' by the
German German(s) may refer to: * Germany, the country of the Germans and German things **Germania (Roman era) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizenship in Germany, see also Ge ...
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Marcus Elieser Bloch Marcus Elieser Bloch (1723–1799) was a German physician and naturalist who is best known for his contribution to ichthyology through his multi-volume catalog of plates illustrating the fishes of the world. Brought up in a Hebrew-speaking Jewish ...
(1723–1799) with the type locality given erroneously as Japan when it was actually Indonesia.


References


External links

* {{Taxonbar, from=Q2791901 boenak Fish of the Indian Ocean Fish of the Pacific Ocean Fish of Thailand Fish described in 1790 Taxa named by Marcus Elieser Bloch Taxonomy articles created by Polbot