''Cephalanthus occidentalis'' is a species of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...

in the family

Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules ...

that is native to eastern and southern North America. Common names include buttonbush, common buttonbush, button-willow, buck brush, and honey-bells.

Description

''Cephalanthus occidentalis'' is a

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...

shrub or small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

that averages in height, but can reach . The leaves are opposite or in whorls of three, elliptic to ovate, long and broad, with a smooth edge and a short petiole. The

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...

s are arranged in a dense spherical

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...

in diameter on a short

peduncle Peduncle may refer to: *Peduncle (botany), a stalk supporting an inflorescence, which is the part of the shoot of seed plants where flowers are formed *Peduncle (anatomy), a stem, through which a mass of tissue is attached to a body **Peduncle (art ...

. Each flower has a fused white to pale yellow four-lobed

corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...

forming a long slender tube connecting to the

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined ...

s. The stigma protrudes slightly from the corolla. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...

is a spherical cluster of

achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not op ...

s (nutlets).

Taxonomy

There are two varieties, not considered distinct by all authorities: *''Cephalanthus occidentalis'' var. ''occidentalis'' (syn. var. ''pubescens'') – common buttonbush. Eastern North America from

Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native En ...

west to

Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the List of U.S. states and territories by population, 22nd most populous, with over 5.75 million residents. Minne ...

and south to

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...

and east

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

. *''Cephalanthus occidentalis'' var. ''californicus'' – California button-willow. Southwestern North America, from west Texas west to

California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...

(

Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...

foothills,

San Joaquin Valley The San Joaquin Valley ( ; es, Valle de San Joaquín) is the area of the Central Valley of the U.S. state of California that lies south of the Sacramento–San Joaquin River Delta and is drained by the San Joaquin River. It comprises seven ...

Sacramento Valley , photo =Sacramento Riverfront.jpg , photo_caption= Sacramento , map_image=Map california central valley.jpg , map_caption= The Central Valley of California , location = California, United States , coordinates = , boundaries = Sierra Nevada (ea ...

, and the

Inner North Coast Ranges The Inner North Coast Ranges occur along the east slope of the outer Northern California Coast Ranges. The term ''inner'' is a reference to the greater distances of the mountain ranges from the Pacific Ocean, compared to the ''outer'' ranges that ...

) and south to

Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...

and

Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...

Habitat

Buttonbush is a common shrub of many

wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...

habitats in its range, including swamps,

floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...

mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...

pocosin A pocosin is a type of palustrine wetland with deep, acidic, sandy, peat soils. Groundwater saturates the soil except during brief seasonal dry spells and during prolonged droughts. Pocosin soils are nutrient-deficient ( oligotrophic), especiall ...

riparian zone A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ...

s, and moist forest

understory In forestry and ecology, understory (American English), or understorey (Commonwealth English The use of the English language in current and former member countries of the Commonwealth of Nations was largely inherited from British co ...

. It is a member of the flora in the

Everglades The Everglades is a natural region A natural region (landscape unit) is a basic geographic unit. Usually, it is a region which is distinguished by its common natural features of geography, geology, and climate. From the ecological point o ...

Ecology

Waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...

and other

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s eat the seeds.

Wood duck The wood duck or Carolina duck (''Aix sponsa'') is a species of perching duck found in North America. The drake wood duck is one of the most colorful North American waterfowl. Description The wood duck is a medium-sized perching duck. A t ...

s utilize the plant as

nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...

protection, and

mallard The mallard () or wild duck (''Anas platyrhynchos'') is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Arge ...

s eat the fruit.

Deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the ...

browse the foliage, which is poisonous to livestock.

Insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...

s and

hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics ar ...

s take the nectar, with

bees Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfam ...

using it to make

honey Honey is a sweet and viscous substance made by several bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primar ...

. It is a larval host to the

hydrangea sphinx ''Darapsa versicolor'', the hydrangea sphinx, is a moth species of the family Sphingidae that inhabits eastern North America, often in wetlands. It was first described by Thaddeus William Harris in 1839. Its wingspan can reach 58 to 80 mm ...

, the

royal walnut moth ''Citheronia regalis'', the regal moth or royal walnut moth, is a North American moth in the family Saturniidae. The caterpillars are called hickory horned devils. The adult ( imago) has a wingspan of . The species was first described by Johan ...

, and the

titan sphinx ''Aellopos titan'', the Titan sphinx, is a moth of the family Sphingidae. The species was first described by Pieter Cramer in 1777. Distribution It has been found all the way from Maine in the United States through Central America and sou ...

Distribution

The species occurs in eastern North America with disjunct populations occurring in the west. In

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...

, it occurs from southern

Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central C ...

and

Quebec Quebec ( ; )According to the Government of Canada, Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is ...

east to

New Brunswick New Brunswick (french: Nouveau-Brunswick, , locally ) is one of the thirteen Provinces and territories of Canada, provinces and territories of Canada. It is one of the three Maritime Canada, Maritime provinces and one of the four Atlantic Canad ...

and south-western

. Besides the eastern United States, and eastern regions of the

Midwest The Midwestern United States, also referred to as the Midwest or the American Midwest, is one of four census regions of the United States Census Bureau (also known as "Region 2"). It occupies the northern central part of the United States. ...

, notable areas range into

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a U.S. state, state in the Southwestern United States. It is the list of U.S. states and territories by area, 6th largest and the list of U.S. states and territories by population, 14 ...

, the

Mogollon Rim The Mogollon Rim ( or or ) is a topographical and geological feature cutting across the northern half of the U.S. state of Arizona. It extends approximately , starting in northern Yavapai County and running eastward, ending near the borde ...

, and other mountain ranges; in California, the entire

West of the Great Plains and the Rocky Mountains, only western Texas, Arizona, and California find ''C. occidentalis''.

Uses

Medicinal

''Cephalanthus occidentalis'' has a number of historical

medicinal Medicine is the science and practice of caring for a patient, managing the diagnosis, prognosis, prevention, treatment, palliation of their injury or disease, and promoting their health. Medicine encompasses a variety of health care practic ...

uses, but it is also

toxic Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subs ...

due to the presence of cephalanthin.

Cultivation

Buttonbush is cultivated as an

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...

for a nectar source or '

honey plant Honeybees usually collect nectar, pollen, or both from the following species of plants, which are called honey plants, for making honey. Acanthaceae (Acanthus family) *'' Avicennia nitida'' Jacq. or Avicennia ...

' and for aesthetics in

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...

s and

native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equ ...

landscapes, and is planted on slopes to help control

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is di ...

. Buttonbush is a suitable shrub for butterfly gardens.

San Joaquin Valley landmark tree

Buttonwillow tree in Buttonwillow, California

The town of

Buttonwillow, California Buttonwillow is an unincorporated community and census-designated place in the San Joaquin Valley, in Kern County, California. Buttonwillow is west of Bakersfield, at an elevation of ). The population was 1,508 at the 2010 census, up from 1,266 ...

was named for the buttonbush (''Cephalanthus occidentalis''). A lone buttonbush served as a landmark on an old trans-San Joaquin Valley trail, and was used by ancient

Yokuts The Yokuts (previously known as MariposasPowell, 1891:90–91.) are an ethnic group of Native Americans native to central California. Before European contact, the Yokuts consisted of up to 60 tribes speaking several related languages. ''Yokut ...

as a meeting place. It later became the site of settlers' stock rodeos. This buttonbush tree is listed as

California Historical Landmark A California Historical Landmark (CHL) is a building, structure, site, or place in California that has been determined to have statewide historical landmark significance. Criteria Historical significance is determined by meeting at least one of ...

No. 492, and is now known as the "Buttonwillow Tree."

References

External links

USDA Plants Profile: ''Cephalanthus occidentalis''

* ttp://ucjeps.berkeley.edu/cgi-bin/get_JM_treatment.pl?Cephalanthus+occidentalis Jepson Flora Project: ''Cephalanthus occidentalis'' var. ''californicus'' {{Taxonbar, from=Q4992358

occidentalis This list of Latin and Greek words commonly used in systematic names is intended to help those unfamiliar with classical languages to understand and remember the scientific names of organisms. The binomial nomenclature used for animals and plants i ...

Flora of North America Medicinal plants of North America Garden plants of North America Plants described in 1753 Taxa named by Carl Linnaeus