Centropus Sinensis
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A coucal is one of about 30
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s in the
cuckoo Cuckoos are birds in the Cuculidae ( ) family, the sole taxon in the order Cuculiformes ( ). The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals, and anis. The coucals and anis are somet ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
. All of them belong in the subfamily Centropodinae and the genus ''Centropus''. Unlike many
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
cuckoos, coucals are not
brood parasite Brood may refer to: Nature * Brood, a collective term for offspring * Brooding, the incubation of bird eggs by their parents * Bee brood, the young of a beehive * Individual broods of North American periodical cicadas: ** Brood X, the largest ...
s, though they do have their own reproductive peculiarity: all members of the genus are (to varying degrees) sex-role reversed, so that the smaller male provides most of the parental care. Male pheasant coucals (''Centropus phasianinus'') invest in building the nest, incubate for the most part and take a major role in feeding the young. At least one coucal species, the
black coucal The black coucal (''Centropus grillii'') is a species of cuckoo in the family Cuculidae. It has a wide distribution in Africa south of the Sahara. Description The male black coucal is in length while the female is , otherwise they are similar i ...
, is
polyandrous Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wives ...
.


Taxonomy

The genus ''Centropus'' was introduced in 1811 by the German zoologist
Johann Karl Wilhelm Illiger Johann Karl Wilhelm Illiger (19 November 1775 – 10 May 1813) was a German entomologist and zoologist. He founded the entomological periodical ''Magazin für Insektenkunde.'' The plant genus '' Illigera'' is named in his honour. Biography Illi ...
. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was subsequently designated as the Senegal coucal by
George Robert Gray George Robert Gray (8 July 1808 – 6 May 1872) was an English zoology, zoologist and author, and head of the Ornithology, ornithological section of the British Museum, now the Natural History Museum, London, Natural History Museum, London f ...
in 1840. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''kentron'' meaning "spur" or "spike" with ''pous'' meaning "foot".


Description

Many coucals have a long claw on their hind toe (hallux). The feet have minute spurs and this is responsible for the German term for coucals ''Sporenkuckucke''. The common name is perhaps derived from the French ''coucou'' and ''alouette'' (for the long lark like claw). (Cuvier, in Newton 1896) The length of the claw can be about 68-76% of the tarsus length in the African black coucal ''C. grillii'' and lesser coucal ''C. bengalensis''. Only the short-toed coucal ''C. rectunguis'' is an exception with the hallux claw of only 23% of the tarsus length. Thread like feather structures (elongated sheaths of the growing feathers that are sometimes termed trichoptiles) are found on the head and neck of hatchlings and can be as long as 20mm. Nestlings can look spiny. Many are opportunistic predators, ''Centropus phasianus'' is known to attack birds caught in mist nets while white-browed coucals ''Centropus superciliosus'' are attracted to smoke from grass fires where they forage for insects and small mammals escaping from the fire. Coucals generally make nests inside dense vegetation and they usually have the top covered but some species have the top open. Pheasant coucal ''Centropus phasianinus'', greater coucal ''C. sinensis'' and Madagascar coucal ''C. toulou'' sometimes build an open nest while some species always build open nests (the bay coucal ''C. celebensis'') Some coucal species have been seen to fly while carrying their young.


Species

The genus contains 29 species: *
Buff-headed coucal The buff-headed coucal (''Centropus milo'') is a species of coucal. These are often placed in the cuckoo family (biology), family (Cuculidae) but seem to warrant recognition as a distinct family. ''C. milo'' is a common Endemism, endemic of the ...
, ''Centropus milo'' * White-necked coucal or pied coucal, ''Centropus ateralbus'' * Ivory-billed coucal or greater black coucal, ''Centropus menbeki'' * Biak coucal, ''Centropus chalybeus'' * Rufous coucal, ''Centropus unirufus'' *
Green-billed coucal The green-billed coucal (''Centropus chlororhynchos'') is a member of the cuckoos. It is endemic to Sri Lanka's wet zone and listed as Vulnerable on the IUCN Red List, as the small population declined due to forest destruction and fragmentation ...
, ''Centropus chlororhynchos'' * Black-faced coucal, ''Centropus melanops'' *
Black-hooded coucal The black-hooded coucal (''Centropus steerii'') is a species of cuckoo in the family Cuculidae. It is endemic to Mindoro in the Philippines and is one of the most endangered birds in the country and is the only critically endangered coucal in the ...
, ''Centropus steerii'' *
Short-toed coucal The short-toed coucal (''Centropus rectunguis'') is a species of cuckoo in the family Cuculidae. It is found in Brunei, Indonesia, Malaysia, and Thailand. Its natural habitats are subtropical or tropical moist lowland forest and subtropical or tr ...
, ''Centropus rectunguis'' * Bay coucal, ''Centropus celebensis'' * Gabon coucal, ''Centropus anselli'' * Black-throated coucal, ''Centropus leucogaster'' * Senegal coucal, ''Centropus senegalensis'' * Blue-headed coucal, ''Centropus monachus'' * Coppery-tailed coucal, ''Centropus cupreicaudus'' * White-browed coucal, ''Centropus superciliosus'' *
Burchell's coucal Burchell's coucal (''Centropus burchellii'') is a species of cuckoo in the family Cuculidae. It is found in the southeastern parts of sub-Saharan Africa. It inhabits areas with thick cover afforded by rank undergrowth and scrub, including in sui ...
, ''Centropus burchellii'' *
Sunda coucal The Sunda coucal (''Centropus nigrorufus''), also known as the Javanese lathe, is a cuckoo species in the family Cuculidae that is endemic to Java, Indonesia. It inhabits mangroves, freshwater swamp forests and grasslands near brackish water. It ...
, ''Centropus nigrorufus'' *
Greater coucal The greater coucal or crow pheasant (''Centropus sinensis''), is a large non-parasitic member of the cuckoo Order (biology), order of birds, the Cuculiformes. A widespread resident in the Indian Subcontinent and Southeast Asia, it is divided into ...
, ''Centropus sinensis'' *
Malagasy coucal The Malagasy coucal or Madagascar coucal (''Centropus toulou'') is a species of cuckoo in the family Cuculidae. It is found in Madagascar and in the Seychelles, where it occurs on Aldabra and was formerly present on Assumption Island and Cosm ...
or Madagascar coucal, ''Centropus toulou'' * Goliath coucal, ''Centropus goliath'' *
Black coucal The black coucal (''Centropus grillii'') is a species of cuckoo in the family Cuculidae. It has a wide distribution in Africa south of the Sahara. Description The male black coucal is in length while the female is , otherwise they are similar i ...
, ''Centropus grillii'' * Philippine coucal, ''Centropus viridis'' *
Lesser coucal The lesser coucal (''Centropus bengalensis'') is a species of cuckoo in the family Cuculidae. It has a wide distribution range that overlaps with several other similar species. The habitat in which it is found is often marshy land with grass and ...
, ''Centropus bengalensis'' *
Violaceous coucal The violaceous coucal or violet coucal (''Centropus violaceus'') is a cuckoo species in the family Cuculidae. It is endemic to the Bismarck Archipelago (Papua New Guinea). Its natural habitat is subtropical or tropical moist lowland forests. Du ...
, ''Centropus violaceus'' * Black-billed coucal or lesser black coucal, ''Centropus bernsteini'' * Kai coucal, ''Centropus spilopterus'' * Pheasant coucal, ''Centropus phasianinus'' * Andaman coucal or brown coucal, ''Centropus andamanensis'' A fossil species, '' Centropus colossus,'' is known from the Quaternary-aged
Fossil Cave Fossil Cave (5L81), formerly known as The Green Waterhole, is a cave in the Limestone Coast region of south-eastern South Australia. It is located in the gazetted locality of Tantanoola, South Australia, Tantanoola about north-west of the c ...
,
Tantanoola, South Australia Tantanoola is a town in regional South Australia. The name is derived from the aboriginal word ''tentunola'', which means ''boxwood / brushwood hill or camp''. ''Tantanoola'' was originally named 'Lucieton' by Governor Jervois after his daughte ...
.


References

{{Taxonbar, from=Q657775 *