Cemophora Coccinea
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''Cemophora coccinea'', commonly known as the scarlet snake, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of nonvenomous snake in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Colubrinae The Colubrinae are a subfamily of snakes within the family Colubridae. It includes numerous genus, genera, and although Taxonomy (biology), taxonomic sources often disagree on the exact number, the Reptile Database lists 717 species in 92 genera ...
of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...
. The species is native to the
southeastern United States The Southeastern United States, also known as the American Southeast or simply the Southeast, is a geographical List of regions in the United States, region of the United States located in the eastern portion of the Southern United States and t ...
. There are two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of ''C. coccinea'' that are recognized as being valid. The Texas scarlet snake (''C. lineri)'' was previously considered a subspecies.


Description

The scarlet snake is relatively small, semi fossorial habitat specialists from southeastern United States, growing to a total length (tail included) of 14–26 inches (36–66 cm) at adult size. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
pattern consists of a light
gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
ground color, with a series of
black Black is a color that results from the absence or complete absorption of visible light. It is an achromatic color, without chroma, like white and grey. It is often used symbolically or figuratively to represent darkness.Eva Heller, ''P ...
-bordered
red Red is the color at the long wavelength end of the visible spectrum of light, next to orange and opposite violet. It has a dominant wavelength of approximately 625–750 nanometres. It is a primary color in the RGB color model and a seconda ...
,
white White is the lightest color and is achromatic (having no chroma). It is the color of objects such as snow, chalk, and milk, and is the opposite of black. White objects fully (or almost fully) reflect and scatter all the visible wa ...
or
yellow Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In t ...
blotches down the back. The belly is either a uniform light gray or white color. The dorsal blotches can extend down the sides of the body, appearing somewhat like banding or rings, which sometimes leads to confusion with other
sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...
species such as the
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
coral snake Coral snakes are a large group of elapid snakes that can be divided into two distinct groups, the Old World coral snakes and New World coral snakes. There are 27 species of Old World coral snakes, in three genera ('' Calliophis'', '' Hemibungar ...
s or the harmless scarlet king snake. The similar pattern exhibited by these three snakes is a form of mimicry known as batesian mimicry. This is a phenomenon in which members of a palatable species gain protection from predation by resembling or mimicking the defensive signaling of an unpalatable or defended species.


Etymology

The generic name, ''Cemophora'', is derived from the Greek words ''Kemos'', meaning "muzzle", and ''phoros'', meaning "bearing", referring to the pronounced rostral scale characteristic of members of the genus.


Geographic distribution

''C. coccinea'' is found only in the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
. It is native to peninsular
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
, and found in southeastern
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
, eastern
Oklahoma Oklahoma ( ; Choctaw language, Choctaw: , ) is a landlocked U.S. state, state in the South Central United States, South Central region of the United States. It borders Texas to the south and west, Kansas to the north, Missouri to the northea ...
,
Arkansas Arkansas ( ) is a landlocked state in the West South Central region of the Southern United States. It borders Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, Texas to the southwest, and Oklahoma ...
, parts of
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
,
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
,
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
,
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
,
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
,
Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...
,
Kentucky Kentucky (, ), officially the Commonwealth of Kentucky, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Illinois, Indiana, and Ohio to the north, West Virginia to the ...
,
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. It borders on Lake Michigan to its northeast, the Mississippi River to its west, and the Wabash River, Wabash and Ohio River, Ohio rivers to its ...
,
Indiana Indiana ( ) is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Lake Michigan to the northwest, Michigan to the north and northeast, Ohio to the east, the Ohio River and Kentucky to the s ...
,
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
,
Maryland Maryland ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic region of the United States. It borders the states of Virginia to its south, West Virginia to its west, Pennsylvania to its north, and Delaware to its east ...
, and
Delaware Delaware ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic and South Atlantic states, South Atlantic regions of the United States. It borders Maryland to its south and west, Pennsylvania to its north, New Jersey ...
; with
disjunct population In biology, a taxon with a disjunct distribution is one that has two or more groups that are related but considerably separated from each other geographically. The causes are varied and might demonstrate either the expansion or contraction of a s ...
s in
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
and central
Missouri Missouri (''see #Etymology and pronunciation, pronunciation'') is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it border ...
. The species is more commonly found throughout most of the
Atlantic coastal plain The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
areas. It prefers open forested areas with sandy soil, ground litter, and organic debris. In
Indiana Indiana ( ) is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Lake Michigan to the northwest, Michigan to the north and northeast, Ohio to the east, the Ohio River and Kentucky to the s ...
, the scarlet snake is listed as an
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
. In New Jersey, the scarlet snake has been recommended by the New Jersey Endangered and Nongame Advisory Committee to be given threatened status for the species within the state, but no formal rule proposal has been filed to date. The threatened status is largely due to population declines and habitat loss. Reasons for the decline in the population are loss of habitat, illegal capture for the pet trade, road mortality, and direct intentional killing.


Behavior and diet

The scarlet snake is
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
and is active only during the summer months. It can be found during the day beneath logs and under pine debris or other organic litter. It also burrows underground during daytime hours, or uses previously dug tunnels as shelter. At night it is often seen crossing roads, or prowling the forest floor searching for prey. The diet of ''C. coccinea'' consists of
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s, small
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s, and
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
of lizards, turtles, and other snakes. Its large, very sharp posterior teeth are used to slash open large reptile eggs. The snake will either squeeze an egg to expel its contents or thrust its head into the egg to break it open. The smallest reptile eggs are eaten in their entirety.


Reproduction

There is very little known about the reproductive habits of the scarlet snake. It is
oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...
, generally laying 2–9 eggs per
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
, with the typical clutch yielding five eggs. Breeding occurs throughout the spring months, and eggs are laid throughout the summer in burrows or under rocks. The eggs hatch two months after being laid, typically in the late summer or autumn. In Florida, one female scarlet snake laid 13 fertile eggs, which exceeded the largest previously reported clutch of 9 eggs.


Predators and defense

The natural predators of the scarlet snake are snake-eating snakes such as
coral snake Coral snakes are a large group of elapid snakes that can be divided into two distinct groups, the Old World coral snakes and New World coral snakes. There are 27 species of Old World coral snakes, in three genera ('' Calliophis'', '' Hemibungar ...
s, and predatory birds and mammals. The scarlet snake rarely bites when picked up by humans, but it can release a foul-smelling odor. The scarlet snake will use its version of batesian mimicry and mimic the venomous coral snakes as a defense mechanism in order to reduce predation.


Conservation

The two greatest threats that scarlet snakes face are the destruction of their habitats because of commercial development and the rising rate of road mortality. Other threats are illegal capture of the species for the pet trade and intentional killing.


Subspecies

The following two subspecies are recognized as being valid. www.reptile-database.org. *'' Cemophora coccinea coccinea'' – Florida scarlet snake *'' Cemophora coccinea copei'' – northern scarlet snake ''
Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a genus other than ''Cemophora''.


References


Further reading

* Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp. . (''Cemophora coccinea'', pp. 592–593 + Plates 595, 596, 607). * Blumenbach F(1788). "''Einige Naturhistorische Bemerkungen bey Gelegenheit einer Schweizer-Reise''". ''Magazin für das Neueste aus der Physik und Naturgeschichte'' 5 (2): 13–23. (''Coluber coccineus'', new species, p. 21). (in German and Latin). * Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition''. The Peterson Field Guide Series. Boston: Houghton Mifflin. xviii + 429 pp. + Plates 1–48. (hardcover); (paperback). (''Cemophora coccinea'', pp. 211–212 + Plates 30, 31 + Map 152). *Conant R, Bridges W (1939). ''What Snake Is That?: A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (with 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century. Frontispiece map + viii + 163 pp. + Plates A–C, 1–32. (''Cemophora coccinea'', pp. 85–86 + Plate 26, Figure 78). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp., 207 Figures, 47 Plates. . (''Cemophora coccinea'', pp. 367–368 + Figure 159 on p. 330 + Plates 32, 44). * Schmidt KP, Davis DD (1941). ''Field Book of Snakes of the United States and Canada''. New York: G.P. Putnam's Sons. 365 pp. (''Cemophora coccinea'', pp. 193–194, Figure 60). * Smith HM, Brodie ED Jr (1982). ''Reptiles of North America: A Guide to Field Identification''. New York: Golden Press. 240 pp. (paperback). (''Cemophora coccinea'', pp. 178–179). * Stejneger L, Barbour T (1917). ''A Check List of North American Amphibians and Reptiles''. Cambridge, Massachusetts: Harvard University Press. 125 pp. (''Cemophora coccinea'', p. 91). * Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock. 1,105 pp. (in two volumes) (''Cemophora coccinea'', pp. 111–115, Figure 36, Map 13). * Zim HS, Smith HM (1956). ''Reptiles and Amphibians: A Guide to Familiar American Species''. A Golden Nature Guide. New York: Simon and Schuster. 160 pp. (Scarlet snake, ''Cemophora doliata'', pp. 100, 156).


External links


South Carolina Snake Species: Scarlet Snake
{{Taxonbar, from=Q1787849 Cemophora Snakes of North America Reptiles described in 1788 Taxa named by Johann Friedrich Blumenbach Fauna of the Southeastern United States