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The Celyphidae, commonly known as beetle flies or beetle-backed flies, are a family of flies (order Diptera). About 115
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in about 9 genera are known chiefly from the Oriental and Afrotropic biogeographic regions with one lineage in the New World.


Description

Celyphidae are small to medium-sized and easily recognised. The scutellum is enlarged, and forming a protective shell over the abdomen, giving them a beetle-like appearance. Also, like many beetles, Celyphidae are often shiny or metallic in color. The head has few bristles. The wings, when at rest, are folded beneath the scutellar "shell". The arista of the antenna is often flattened and leaf-like at the base. The family name is derived from the Greek word κέλυφος for pod or shell. Male celyphids lack an aedeagus and instead have paired gonapophyses that are used in copulation and are of key taxonomic value.


Biology

The biology of the family is poorly known. Adults are found along streams and rivers, and in wet, grassy areas. Larvae are saprophagous.


Relatives

The family Celyphidae is considered by most authors to be the sister taxon of the Lauxaniidae (e.g., Griffiths 1972), and this has been supported by some molecular studies which suggest the (Chamaemyiidae + (Lauxaniidae + Celyphidae)) within the Lauxanoiodea. In the past they have occasionally been considered a specialized lineage within the Lauxaniidae.


Classification

*Genus '' Acelyphus'' Malloch, 1929 - Asia *Genus '' Atopocelyphus,'' Gaimari, 2017 - French Guiana *Genus '' Celyphus'' Dalman, 1818 - Africa and Asia **Subgenus '' Celyphus'' Dalman, 1818 - Asia **Subgenus '' Hemiglobus'' Frey, 1941 - Africa **Subgenus '' Paracelyphus'' Bigot, 1859 - Asia *Genus '' Chamaecelyphus'' Frey, 1941 - Africa *Genus '' Idiocelyphus'' Malloch, 1929 - Asia *Genus '' Oocelyphus'' Chen, 1949 - southern China *Genus '' Spaniocelyphus'' Hendel, 1914 - Africa and Asia


References


Further reading

* Keys genera, species. * Keys genera, species. * *Paap, L. Celyphidae in Soós, Á, Paap, L. (Eds.) 1984 - 1992. ''Catalogue of Palaearctic Diptera'' 9 . Akadémiai Kiadó, Budapest - Elsevier, Amsterdam: 63-66..


External links


Research project on CelyphidaeImages from Diptera.infoFamily Celyphidae at EOL
Images from EOL * World list {{Taxonbar, from=Q4583736 Brachycera families