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''Cedrus'', with the common English name cedar, is a genus of
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
ous
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s in the plant family
Pinaceae The Pinaceae (), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, Pinyon_pine, piñons, larches, pines and spruces. The family is incl ...
(subfamily Abietoideae). They are native to the mountains of the western
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
and the Mediterranean region, occurring at altitudes of in the Himalayas and in the Mediterranean.Farjon, A. (1990). ''Pinaceae: Drawings and Descriptions of the Genera''. Koeltz Scientific Books. .


Description

''Cedrus'' trees can grow up to 30–40 m (occasionally 60 m) tall with spicy-resinous scented
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
, thick ridged or square-cracked bark, and broad, level branches. The shoots are dimorphic and are made up of long shoots, which form the framework of the branches, and short shoots, which carry most of the leaves. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
and needle-like, 8–60 mm long, arranged in an open spiral
phyllotaxis In botany, phyllotaxis () or phyllotaxy is the arrangement of leaf, leaves on a plant stem. Phyllotactic spirals form a distinctive class of patterns in nature. Leaf arrangement The basic leaf#Arrangement on the stem, arrangements of leaves ...
on long shoots, and in dense spiral clusters of 15–45 together on short shoots; they vary from bright grass-green to dark green to strongly glaucous pale blue-green, depending on the thickness of the white wax layer which protects the leaves from desiccation. The seed cones are barrel-shaped, 6–12 cm long and 3–8 cm broad, green maturing grey-brown, and, as in ''Abies'', disintegrate at maturity to release the winged
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s. The seeds are 10–15 mm long, with a 20–30 mm wing; as in ''Abies'', the seeds have two or three resin blisters, containing an unpleasant-tasting
resin A resin is a solid or highly viscous liquid that can be converted into a polymer. Resins may be biological or synthetic in origin, but are typically harvested from plants. Resins are mixtures of organic compounds, predominantly terpenes. Commo ...
, thought to be a defence against squirrel predation. Cone maturation takes one year, with pollination in autumn and the seeds maturing at the same time a year later. The
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
cones are slender ovoid, 3–8 cm long, produced in late summer, and shed pollen in autumn.Frankis, M. & Lauria, F. (1994). The maturation and dispersal of cedar cones and seeds. ''International Dendrology Society Yearbook'' 1993: 43–46.


Classification

Cedars share a very similar cone structure with the firs (''Abies'') and were traditionally thought to be most closely related to them, but molecular evidence supports a basal position in the family.Liston A., D.S. Gernandt, T.F. Vining, C.S. Campbell, D. Piñero. 2003. "Molecular Phylogeny of Pinaceae and Pinus". In Mill, R. R. (ed.): ''Proceedings of the 4th Conifer Congress. Acta Hort'' 615: pp. 107–114.Wang, X.-Q., Tank, D. C. and Sang, T. (2000): "Phylogeny and Divergence Times in Pinaceae: Evidence from Three Genomes". ''Molecular Biology and Evolution'' 17:773-781. Availabl
online


Taxonomy

The five
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
of ''Cedrus'' are assigned according to taxonomic opinion to between one and four species:Farjon, A. (2008). ''A Natural History of Conifers''. Timber Press . The oldest known fossil of ''Cedrus'' is ''Cedrus penzhinaensis'' known from fossil wood found in Early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
(
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) sediments of Kamchatka, Russia.


Ecology

Cedars are adapted to mountainous climates; in the Mediterranean, they receive winter precipitation, mainly as snow, and summer drought, while in the western Himalaya, they receive primarily summer monsoon rainfall and occasional winter snowfall. While no members of ''Cedrus'' are native to the Americas, members of ''Juniperus'' and Cupressaceae are native and are called by the common name of "cedar". Cedars are used as food plants by the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of some
Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...
species including pine processionary and turnip moth (recorded on deodar cedar).


Use

Cedars are very popular ornamental trees, and are often cultivated in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
s where winter temperatures do not fall below circa −25 °C. The Turkish cedar is slightly hardier, to −30 °C or just below. Extensive mortality of planted specimens can occur in severe winters when temperatures fall lower.Ødum, S. (1985). "Report on frost damage to trees in Denmark after the severe 1981/82 and 1984/85 winters". Hørsholm Arboretum, Denmark. Locales with successful longaeval cultivation include the Mediterranean region, Western Europe north to the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
, southern
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
and
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
, and southern and western
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Cedar wood and cedarwood oil are natural repellents to moths, hence cedar is a popular lining for cedar chests and closets in which woolens are stored. This specific use of cedar is mentioned in '' The Iliad'', Book 24, referring to the cedar-roofed or lined storage chamber where Priam went to fetch treasures to be used as ransom. The ancients made cedarwood oil from Lebanon cedar, a true cedar of the genus ''Cedrus'', However, the species used for modern cedar chests and closets in North America is '' Juniperus virginiana'', and cedarwood oil is now typically derived from various junipers and cypresses (of the family '' Cupressaceae''). Cedar is also commonly used to make shoe trees because it can absorb moisture and deodorize. Many species of cedar are suitable for training as
bonsai Bonsai (; , ) is the Japanese art of Horticulture, growing and shaping miniature trees in containers, with a long documented history of influences and native Japanese development over a thousand years, and with unique aesthetics, cultural hist ...
. They work well for many styles, including formal and informal upright, slanting, and cascading.


Nomenclature

Some authorities consider ''Cedrus'' the only "true cedars" and discourage use of the name for other genera without an additional qualifier, such as "white-cedar".Kelsey, H. P., & Dayton, W. A. (1942). ''Standardized Plant Names'', second edition. American Joint Committee on Horticultural Nomenclature. Horace McFarland Company, Harrisburg, Pennsylvania. Nevertheless, the name "cedar" has been applied (since about 1700) to other trees, such as the North American ''
Thuja plicata ''Thuja plicata'' is a large evergreen coniferous tree in the family Cupressaceae, native to the Pacific Northwest of North America. Its common name is western redcedar in the U.S. or western red cedar in the UK, and it is also called pacific re ...
'', commonly called "western red cedar", and '' Juniperus virginiana'', commonly called "red cedar" or "eastern red cedar". In some cases, the botanical name alludes to this usage, such as the genus '' Calocedrus'', meaning "beautiful cedar" (also known as "incense cedar"). Several species of genera '' Calocedrus'', '' Thuja'', and '' Chamaecyparis'' in the
Pacific Northwest The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
having similarly aromatic wood are referred to as "false cedars" . In Australia '' Toona ciliata'' has long been known as red cedar, in furniture and as the tree, even though it belongs to the Meliaceae or Mahogany family.


Etymology

Both the Latin word ''cedrus'' and the generic name ''cedrus'' are derived from Greek κέδρος ''kédros''. Ancient Greek and Latin used the same word, ''kédros'' and ''cedrus'', respectively, for different species of plants now classified in the genera ''Cedrus'' and ''Juniperus'' ( juniper). Species of both genera are native to the area where Greek language and culture originated, though as the word ''kédros'' does not seem to be derived from any of the languages of the
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
, it has been suggested the word may originally have applied to Greek species of juniper and was later adopted for species now classified in the genus ''Cedrus'' because of the similarity of their aromatic woods.Meiggs, R. 1982. Trees and Timber in the Ancient Mediterranean World. The name was similarly applied to citron and the word
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
is derived from the same root.Andrews, A. C. 1961. Acclimatization of citrus fruits in the Mediterranean region. ''Agricultural History'' 35: 35–46. However, as a loan word in English, cedar had become fixed to its
biblical The Bible is a collection of religious texts that are central to Christianity and Judaism, and esteemed in other Abrahamic religions such as Islam. The Bible is an anthology (a compilation of texts of a variety of forms) biblical languages ...
sense of ''Cedrus'' by the time of its first recorded usage in AD 1000.''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the principal historical dictionary of the English language, published by Oxford University Press (OUP), a University of Oxford publishing house. The dictionary, which published its first editio ...
''.


See also

* List of plants known as cedar * Cedars of God in the Kadisha Valley of Bsharri,
Lebanon Lebanon, officially the Republic of Lebanon, is a country in the Levant region of West Asia. Situated at the crossroads of the Mediterranean Basin and the Arabian Peninsula, it is bordered by Syria to the north and east, Israel to the south ...


References


External links


Cedar of Meudon
A tree in Meudon near Paris that has fascinated people through time {{Authority control Conifer genera Ornamental trees Plants used in bonsai Wood