''Meta menardi'', the European cave spider, is a long-jawed
orb-weaving spider
Orb-weaver spiders are members of the spider family Araneidae. They are the most common group of builders of spiral wheel-shaped webs often found in gardens, fields, and forests. The English word "orb" can mean "circular", hence the English name ...
(family
Tetragnathidae
Long-jawed orb weavers or long jawed spiders (Tetragnathidae) are a family of araneomorph spiders first described by Anton Menge in 1866. They have elongated bodies, legs, and chelicerae, and build small orb webs with an open hub with few, wid ...
). It is also known as the orbweaving cave spider, the cave orbweaver and, in context, simply the cave spider.
Habitat
They have a natural
range
Range may refer to:
Geography
* Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra)
** Mountain range, a group of mountains bordered by lowlands
* Range, a term used to i ...
extending from
Scandinavia
Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and Swe ...
to
North Africa
North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in t ...
and from
Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
to
Korea
Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republi ...
; there are also transplanted populations as far apart as
Japan and
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
.
The adult spiders are
photophobic
Photophobia is a medical symptom of abnormal intolerance to visual perception of light. As a medical symptom photophobia is not a morbid fear or phobia, but an experience of discomfort or pain to the eyes due to light exposure or by presence of ...
and live in places free from light, frequently in
cave
A cave or cavern is a natural void in the ground, specifically a space large enough for a human to enter. Caves often form by the weathering of rock and often extend deep underground. The word ''cave'' can refer to smaller openings such as sea ...
s and
tunnel
A tunnel is an underground passageway, dug through surrounding soil, earth or rock, and enclosed except for the entrance and exit, commonly at each end. A pipeline is not a tunnel, though some recent tunnels have used immersed tube cons ...
s, though they can sometimes be seen outside of caves and mines as they will emerge around dusk to hunt, often using a single silk lasso line and swinging down upon their prey. They are often found in areas that are frequented by bats. The spiders are most often observed in railway tunnels and mines since these are more likely to be visited by humans. The young spiders are, after several
instar
An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow o ...
s (and in contrast to the adults),
strongly attracted to light—probably an
evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
ary
adaptation which ensures the spread of the species to new areas (see Life cycle for further details).
Morphology
They are shiny satin black to reddish brown in colour, often having a different-coloured rear body which varies from black, brown or even olive green and are one of the largest spiders found in the United Kingdom, adults reaching roughly 5 cm legspan and 15 mm body length. However, the
giant house spider
The giant house spider has been treated as either one species, under the name ''Eratigena atrica'', or as three species, ''E. atrica'', '' E. duellica'' and '' E. saeva''. , the three species view was accepted by the World Spider Catalog. T ...
,
cardinal spider
''Tegenaria parietina'' is a rather rare spider in Europe, with a distribution also including Northern Africa to Central Asia and Sri Lanka, and from the West Indies to Uruguay and Argentina, where it may have been introduced. In the UK it is som ...
and the
raft spider
The raft spider, scientific name ''Dolomedes fimbriatus'', is a large semi-aquatic spider of the family Pisauridae found throughout north-western and central Europe. It is one of only two species of the genus '' Dolomedes'' found in Europe, the o ...
are larger. It is widespread and locally
abundant, although rarely noticed due to its
habits
A habit (or wont as a humorous and formal term) is a routine of behavior that is repeated regularly and tends to occur subconsciously. . They are neither endangered nor protected in the UK.
closely resembles , which sports a fainter mark on its abdomen and no rings on its legs.
The ecology of the two species is very similar. On the other hand, the two species seems to exhibit different tolerance to the microclimatic variations within the cave, which emerged as the main factors determining the differentiation of their niche.
Cave spiders feed on smaller
invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s, most frequently
myriapod
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial.
The fossil record of myriapods reaches back into the late Silurian, a ...
s and
slugs.
The European cave spider is not
dangerous to humans and although (in common with nearly all spiders) they have venom, its effect on large mammals is negligible. They are unlikely to bite if carefully handled but if sufficiently provoked they are capable of inflicting an unpleasant "nip". They are non-aggressive and generally slow moving and are regarded (by
arachnophiles) as "gentle giants".
Life cycle
After mating, females produce a drop-shaped
egg sac
Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species dive ...
(cocoon).
Egg sac
Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species dive ...
s are tear-shaped white pendants (sometimes with a slightly yellow centre) usually hanging from the roof of the habitat by a
silk
Silk is a natural protein fiber, some forms of which can be woven into textiles. The protein fiber of silk is composed mainly of fibroin and is produced by certain insect larvae to form cocoons. The best-known silk is obtained from the ...
thread approximately 20 mm long. Egg sacs are usually laid in the vicinity of the cave entrance, where the air flow velocity ranges between 0.3 and 0.6 m/s.
The cocoon (2–3 cm in radius) contains an average of 200/300 eggs and is laid near the cave entrance at the end of the summer. After hatching, the spiderlings remain in the cocoon until the first molt, feeding on the yolk. They leave the cocoon in spring, move towards the cave entrance and disperse via ballooning outside the cave.
Media mistakes
A
media frenzy
Media circus is a colloquial metaphor, or idiom, describing a news event for which the level of media coverage—measured by such factors as the number of reporters at the scene and the amount of material broadcast or published—is perceived to ...
was caused when
British Telecom
BT Group plc (trade name, trading as BT and formerly British Telecom) is a British Multinational corporation, multinational telecommunications holding company headquartered in London, England. It has operations in around 180 countries and is th ...
engineers working in the grounds of
Windsor Castle
Windsor Castle is a royal residence at Windsor in the English county of Berkshire. It is strongly associated with the English and succeeding British royal family, and embodies almost a millennium of architectural history.
The original c ...
discovered a colony living in
conduit
Conduit may refer to:
Engineering systems
* Conduit (fluid conveyance), a pipe suitable for carrying either open-channel or pressurized liquids
* Electrical conduit, a protective cover, tube or piping system for electric cables
* Conduit curre ...
s and
manhole
A manhole (utility hole, maintenance hole, or sewer hole) is an opening to a confined space such as a shaft, utility vault, or large vessel. Manholes are often used as an access point for an underground public utility, allowing inspection, ...
s. These were incorrectly described as swarms of aggressive spiders, up to 9 cm in size, which a leading entomologist believed were venomous. The
BBC #REDIRECT BBC
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reported that it "may be a new species or a type of spider previously thought to have been extinct for thousands of years."
The initial incident was widely reported but when the species was properly identified the matter received little publicity.
References
Further reading
*
External links
Pictures of ''Meta menardi''at Nick's Spiders
living in Norway
{{Taxonbar, from=Q1470676
Tetragnathidae
Spiders of Europe
Spiders of Africa
Spiders of Asia
Cave spiders
Spiders described in 1804