The caudofemoralis (from the Latin ''cauda'', tail and ''femur'', thighbone) is a
muscle
Muscle is a soft tissue, one of the four basic types of animal tissue. There are three types of muscle tissue in vertebrates: skeletal muscle, cardiac muscle, and smooth muscle. Muscle tissue gives skeletal muscles the ability to muscle contra ...
found in the pelvic limb of mostly all animals possessing a tail. It is thus found in nearly all
tetrapod
A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s.
Location
The caudofemoralis spans
plesiomorphic
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades.
Plesiomorphy, symplesiomorphy, apomorphy, an ...
ally between femur (thigh) and tail; in mammals it is reduced and found directly
posterior/
caudal to the
gluteus maximus
The gluteus maximus is the main extensor muscle of the hip in humans. It is the largest and outermost of the three gluteal muscles and makes up a large part of the shape and appearance of each side of the hips. It is the single largest muscle in ...
and directly
anterior
Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
/
cranial
Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
to the
biceps femoris
The biceps femoris () is a muscle of the thigh located to the posterior, or back. As its name implies, it consists of two heads; the long head is considered part of the hamstring muscle group, while the short head is sometimes excluded from this ...
.
Origin and insertion
The caudofemoralis originates from the transverse processes of the second, third and fourth caudal vertebrae
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
.
The caudodistal portion of the muscle in mammals lies deep to the proximocranial portion of the Biceps femoris; near the middle of the thigh
In anatomy, the thigh is the area between the hip (pelvis) and the knee. Anatomically, it is part of the lower limb.
The single bone in the thigh is called the femur. This bone is very thick and strong (due to the high proportion of bone tissu ...
, the caudofemoralis gives rise to a long, thin, and narrow tendon
A tendon or sinew is a tough band of fibrous connective tissue, dense fibrous connective tissue that connects skeletal muscle, muscle to bone. It sends the mechanical forces of muscle contraction to the skeletal system, while withstanding tensi ...
that passes distally to the knee joint and inserts into the fascia lata that is anchored to the lateral border of the patella
The patella (: patellae or patellas), also known as the kneecap, is a flat, rounded triangular bone which articulates with the femur (thigh bone) and covers and protects the anterior articular surface of the knee joint. The patella is found in m ...
.[
]
Among
archosaurians, the caudofemoralis is divided in a ''pars pelvica/brevis'' (characterized by a pelvic origin) and a ''pars caudalis/longa'' (caudal origin), and the insertion on the femur is marked by the
fourth trochanter (but this becomes reduced in
maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, ...
ns and absent in
birds
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
).
Action
In mammals the caudofemoralis acts to flex the tail laterally to its corresponding side when the pelvic limb is bearing weight. When the pelvic limb is lifted off the ground, contraction of the caudofemoralis causes the limb to abduct and the shank to extend by extending the
hip joint (acetabulofemoral or coxofemoral joint). In other tetrapods contraction of the caudofemoralis retracts the hindlimb.
Cited References
Vertebrate anatomy
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