Caucasian Squirrel
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The Caucasian squirrel (''Sciurus anomalus'') or Persian squirrel, is a
tree squirrel Tree squirrels are the members of the squirrel Family (biology), family (Sciuridae) commonly just referred to as "squirrels". They include more than 100 arboreal species native to all continents except Antarctica and Oceania. They do not form a ...
in the genus ''
Sciurus The genus ''Sciurus'' ( or ) contains most of the common, bushy-tailed squirrels in North America, Europe, Temperate zone, temperate Asia, Central America and South America. Species The number of species in the genus is subject to change. In 200 ...
'' found in
temperate broadleaf and mixed forests Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. These ...
in south-western Asia. The species is traditionally said to have first been described in 1778 by
Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in the 13th edition of ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'', and named ''Sciurus anomalus''. However, this work was actually published in 1788, so the true first description was made by
Johann Anton Güldenstädt Johann Anton Güldenstädt (26 April 1745 in Riga, Latvia – 23 March 1781 in St. Petersburg, Russia) was a Baltic German natural history, naturalist and explorer in Russian service. Güldenstädt lost both his parents early, and from 1763 on ...
in 1785. Gmelin made reference to this in the 1788 work, citing Güldenstädt as the
author In legal discourse, an author is the creator of an original work that has been published, whether that work exists in written, graphic, visual, or recorded form. The act of creating such a work is referred to as authorship. Therefore, a sculpt ...
.


Description

Caucasian squirrels are small tree squirrels, with a total length of , including the tail, and weighing . The color of the upper body fur ranges from greyish brown to pale grey, depending on the subspecies, while that of the underparts is rusty brown to yellowish, and that of the tail, yellow brown to deep red. The claws are relatively short, compared with those of other tree squirrels, and females have either eight or ten teats.
Samuel Griswold Goodrich Samuel Griswold Goodrich (August 19, 1793 – May 9, 1860), better known by his pseudonym Peter Parley, was an American author and politician who established the children's magazine ''Merry's Museum''. He was a Massachusetts Senate, Massachuse ...
described the Caucasian squirrel in 1885 as "Its color is grayish-brown above, and yellowish-brown below".


Distribution and habitat

Caucasian squirrels are native to south-western Asia, where they are found from Turkey, and the islands of
Gökçeada Imbros (; ; ), officially Gökçeada () since 29 July 1970,Alexis Alexandris, "The Identity Issue of The Minorities in Greece And Turkey", in Hirschon, Renée (ed.), ''Crossing the Aegean: An Appraisal of the 1923 Compulsory Population Exchang ...
and
Lesbos Lesbos or Lesvos ( ) is a Greek island located in the northeastern Aegean Sea. It has an area of , with approximately of coastline, making it the third largest island in Greece and the List of islands in the Mediterranean#By area, eighth largest ...
in the west,
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
in the southeast, and as far as
Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...
and
Jordan Jordan, officially the Hashemite Kingdom of Jordan, is a country in the Southern Levant region of West Asia. Jordan is bordered by Syria to the north, Iraq to the east, Saudi Arabia to the south, and Israel and the occupied Palestinian ter ...
in the south. It is one of only two species of the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Sciurus The genus ''Sciurus'' ( or ) contains most of the common, bushy-tailed squirrels in North America, Europe, Temperate zone, temperate Asia, Central America and South America. Species The number of species in the genus is subject to change. In 200 ...
'' to be found on Mediterranean islands, and, although Eurasian red squirrels have been recently introduced to some areas, is the only species of ''Sciurus'' native to the wider region. The species mainly lives in forested areas dominated by
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
,
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
, and
pistachio The pistachio (, ; ''Pistacia vera'') is a small to medium-sized tree of the Anacardiaceae, cashew family, originating in Iran. The tree produces nut (fruit)#Culinary definition and uses, seeds that are widely consumed as food. In 2022, world ...
, up to altitudes of . Three subspecies are recognised: * ''S. a. anomalus'' - Turkey and the Caucasus * ''S. a. pallescens'' -
Zagros Mountains The Zagros Mountains are a mountain range in Iran, northern Iraq, and southeastern Turkey. The mountain range has a total length of . The Zagros range begins in northwestern Iran and roughly follows Iran's western border while covering much of s ...
, from south-eastern Turkey to Iran * ''S. a. syriacus'' -
Lebanon Lebanon, officially the Republic of Lebanon, is a country in the Levant region of West Asia. Situated at the crossroads of the Mediterranean Basin and the Arabian Peninsula, it is bordered by Syria to the north and east, Israel to the south ...
,
Syria Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...
, Israel, and Jordan


Biology and behavior

The squirrels are diurnal, and solitary, although temporary groups may forage where food is plentiful. Their diet includes nuts, seeds, tree shoots, and buds, with the seeds of oak and pine being particularly favored. Like many other squirrels, they cache their food within tree cavities or loose soil, with some larders containing up to of seeds. They live in trees, where they make their dens, but frequently forage on the ground, and are considered less
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
than Eurasian red squirrels. They commonly nest in tree hollows lined with moss and leaves, and located above the ground, but nests are also sometimes found under rocks or tree roots. Their alarm call is high-pitched, and said to resemble the call of the European green woodpecker, and they mark their territories with urine and dung. Breeding occurs throughout the year, but is more common in spring or autumn. Litters range from two to seven, with three or four being typical, and the young are fully mature by five or six months of age.


Conservation

A survey in 2008 found that the species remained abundant within Turkey, however declines are noted in population within the
Levant The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
region. The guides for a survey in 1993 in Israel stated that they considered the species to be nearly extinct within the area studied. Whilst the Caucasian squirrel is threatened by poaching and deforestation, the declines recorded are not sufficient to qualify them as anything other than "Least Concern" by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
. Hunting of the species is banned by the Central Hunting Commission, and the Caucasian squirrel is protected by the Bern Convention and the EU Habitats Directive.


References


External links

{{Taxonbar, from=Q756912 Caucasian squirrel Mammals of West Asia Mammals of Azerbaijan Mammals of Turkey Caucasian squirrel Caucasian squirrel Taxonomy articles created by Polbot Habitats Directive species Taxa named by Johann Anton Güldenstädt