Pinaceae
The Pinaceae (), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, Pinyon_pine, piñons,
larches, pines and spruces. The family is incl ...
, with one known living species, ''Cathaya argyrophylla''. In foliage and cone morphology, ''Cathaya'' has been considered a member of the subfamily Laricoideae, closely related to '' Pseudotsuga'' and '' Larix'', but more recent genetic studies have suggested a closer relationship to ''
Pinus
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae.
''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as c ...
'' and ''
Picea
A spruce is a tree of the genus ''Picea'' ( ), a genus of about 40 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Northern hemisphere. ''Picea'' is the sole genus ...
'' in the subfamily Pinoideae. A second species, ''C. nanchuanensis'', is now treated as a
synonym
A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
, as it does not differ from ''C. argyrophylla'' in any characters.
Description
''Cathaya'' is confined to a limited area in southern
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, in the provinces of
Guangxi
Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...
,
Guizhou
)
, image_skyline =
, image_caption =
, image_map = Guizhou in China (+all claims hatched).svg
, mapsize = 275px
, map_alt = Map showing the location of Guizhou Province
, map_caption = Map s ...
,
Hunan
Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
and southeast
Sichuan
Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
. It is an evergreen
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
growing to tall and a trunk up to diameter, found on steep, narrow
mountain
A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited summit area, and is usually higher t ...
slopes at altitude on
limestone
Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
soils. A larger population has been reduced by over-cutting before its scientific discovery and protection in 1950.
The
leaves
A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are needle-like, long, have ciliate (hairy) margins when young, and grow around the stems in a spiral pattern. The cones are long, with about 15–20 scales, each scale bearing two winged
seed
In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s.
One or two botanists, unhappy with the idea of a new genus in such a familiar family, tried to shoehorn it into other existing genera, as ''Pseudotsuga argyrophylla'' and ''
Tsuga
''Tsuga'' (, from Japanese (), the name of '' Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The English-language common name "hemlock" arose from a perceived similarity in the smell of it ...
argyrophylla''. It is however very distinct from both of these genera, and these combinations are not now used.
The species was introduced into Europe and North America in the 1990s.
Fossil record
The extinct
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
species ''Cathaya loehri'' (Engelhardt & Kinkelin) Chun & Kuang is described from the
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
and
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58 other ''Cathaya'' fossils are known from the Miocene and Pliocene of eastern Siberia, and from the early
Pleistocene
The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
of southern
Portugal
Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
.Forest Context and Policies in Portugal: Present and Future Challenges by Fernando Reboredo – Springer, 28. aug. 2014 – They are abundant in European
brown coal
Lignite (derived from Latin ''lignum'' meaning 'wood'), often referred to as brown coal, is a soft, brown, Combustion, combustible sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35% and is considered ...
deposits dating from between 10 and 30 million years ago.