The Caspian whipsnake (''Dolichophis caspius'', sometimes also ''Coluber caspius'') also known as the large whipsnake (among various other species in genus ''Dolichophis''/''
Coluber''), is a common species of
whipsnake found in the
Balkans
The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
and parts of
Eastern Europe
Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural and socio-economic connotations. Its eastern boundary is marked by the Ural Mountain ...
.
Description
The Caspian whipsnake is perhaps the largest species of snake in Europe. It typically grows to around in length, though a few may exceed a length of . The record sized specimen was approximately
Body mass can be commonly from . This species is not venomous and is mostly
active during the day. The head is joined to the body by a thick neck. Its pupils are round. Nineteen rows of smooth
scales can be found at the middle body, though rarely there may be seventeen. Dorsal scales each have two apical pits at the posterior edge. The centre of the scale is lighter in colour than the edges.
The
dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
The fus ...
side is grey-brown and features markings which are distinctive in juveniles and young snakes but fade with age. The ventral side is light yellow or white.
Taxonomy
Synonyms
Since its original description by
Johann Friedrich Gmelin
Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist.
Education
Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in 1789, ''D. caspius'' has been known by many names. A list of such
synonym
A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
s is given below:
*''Coluber caspius''
*''Coluber acontistes''
*''Coluber thermalis''
*''Coluber jugularis caspius''
*''Hierophis caspius''
*''Dolichophis caspius''
''
Nota bene
( ; plural: ) is the Latin language, Latin phrase meaning ''note well''.
In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': In a synonymy, names without dashes are new
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
(described as new species). However, names with dashes are just new combinations (moved to a different genus, changed from species to subspecies, etc.).
Distribution
The Caspian whipsnake is found in the
Balkan peninsula
The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
, parts of Eastern Europe and a small portion of Asia Minor. It may be found in
Albania
Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
,
Bulgaria
Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
,
Greece
Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
,
North Macedonia
North Macedonia, officially the Republic of North Macedonia, is a landlocked country in Southeast Europe. It shares land borders with Greece to the south, Albania to the west, Bulgaria to the east, Kosovo to the northwest and Serbia to the n ...
,
Serbia
, image_flag = Flag of Serbia.svg
, national_motto =
, image_coat = Coat of arms of Serbia.svg
, national_anthem = ()
, image_map =
, map_caption = Location of Serbia (gree ...
,
Romania
Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
,
Croatia
Croatia, officially the Republic of Croatia, is a country in Central Europe, Central and Southeast Europe, on the coast of the Adriatic Sea. It borders Slovenia to the northwest, Hungary to the northeast, Serbia to the east, Bosnia and Herze ...
,
Turkey
Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
,
Moldova
Moldova, officially the Republic of Moldova, is a Landlocked country, landlocked country in Eastern Europe, with an area of and population of 2.42 million. Moldova is bordered by Romania to the west and Ukraine to the north, east, and south. ...
,
Montenegro
, image_flag = Flag of Montenegro.svg
, image_coat = Coat of arms of Montenegro.svg
, coa_size = 80
, national_motto =
, national_anthem = ()
, image_map = Europe-Mont ...
, southern
Ukraine
Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...
, southern
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
,
Kazakhstan
Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
, and southern
Hungary
Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
.
Results from a survey indicate their habitats may be distributed along low areas near major rivers, such as the
Danube
The Danube ( ; see also #Names and etymology, other names) is the List of rivers of Europe#Longest rivers, second-longest river in Europe, after the Volga in Russia. It flows through Central and Southeastern Europe, from the Black Forest sou ...
and the
Olt River
The Olt ( Romanian and Hungarian; ; or ', , ''Alytos'') is a river in Romania. It is long, and its basin area is . It is the longest river flowing exclusively through Romania. Its average discharge at the mouth is . It originates in the Hă ...
.
It was previously assumed extinct in
Moldavia
Moldavia (, or ; in Romanian Cyrillic alphabet, Romanian Cyrillic: or ) is a historical region and former principality in Eastern Europe, corresponding to the territory between the Eastern Carpathians and the Dniester River. An initially in ...
(eastern Romania, southern Ukraine, and western Moldova), where it was only known from two sites, and not observed since 1937.
Three specimens were collected in May 2007 in
Galați County
Galați () is a county (județ) of Romania, in Moldavia region, with the capital city at Galați, between latitude, 45°25'N and latitude, 46°10'N latitude, longitude, 27°20'E and longitude, 28°10'E longitude. It borders the counties of Vaslu ...
, somewhat dispelling this belief.
Though it is known to be common in the
Dobrudja
Dobruja or Dobrudja (; or ''Dobrudža''; , or ; ; Dobrujan Tatar: ''Tomrîğa''; Ukrainian language, Ukrainian and ) is a Geography, geographical and historical region in Southeastern Europe that has been divided since the 19th century betw ...
area, knowledge of its distribution in other areas is poor. It is believed to be very rare in these parts, and may also be highly threatened. National legislation has declared ''D. caspius'' to be "a species of community interest" and thus must be strictly protected.
In Hungary, it was formerly thought only to exist in two areas: one near the capital of
Budapest
Budapest is the Capital city, capital and List of cities and towns of Hungary, most populous city of Hungary. It is the List of cities in the European Union by population within city limits, tenth-largest city in the European Union by popul ...
, and another near
Szársomlyó. However, a recent survey of the region has yielded several previously unknown whipsnake habitats along the River Danube, including sites near
Dunaújváros
Dunaújváros (; also known by #Etymology and names, alternative names) is an industrial city in Fejér County, Central Hungary. It is a city with county rights. Situated 70 kilometres (43 miles) south of Budapest on the Danube, the city is best ...
and
Dunaföldvár.
As in Romania, ''D. caspius'' is a protected species in Hungary.
Habitat
Recently observed specimens of ''D. caspius'' were found in grassy, open areas with sparse
shrub
A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
bery. They have also been found in open, bushy areas, specifically, between the
Danube Meadow and
Romanian Plain, near
Căscioarele.
Vine covered areas were found near this habitat as well. At
Drăgăneşti Olt, the species was found on a hilly slope.
Its habitat also includes rocky riverbanks and swampy areas, as well as gardens and cemeteries.
Diet
The Caspian whipsnake is a generalist predator and feeds on various vertebrates. Its diet may include small mammals, lizards, birds, and other snakes.
Though not venomous, the Caspian whipsnake is considered "ferocious" and bites quickly and without warning. Due to its ability to consume rodents which may harm crops, it is valued as natural
pest control
Pest control is the regulation or management of a species defined as a pest (organism), pest; such as any animal, plant or fungus that impacts adversely on human activities or environment. The human response depends on the importance of the da ...
.
Conservation
Data on the conservation status of ''D. caspius'' in many regions are sparse. Whilst it is known to be common in the
Dobrudja
Dobruja or Dobrudja (; or ''Dobrudža''; , or ; ; Dobrujan Tatar: ''Tomrîğa''; Ukrainian language, Ukrainian and ) is a Geography, geographical and historical region in Southeastern Europe that has been divided since the 19th century betw ...
area, it may be rare and threatened in other areas. In
Drăgăneşti Olt, Romania,
road-killed snakes are a "common sight" to locals. Traffic-related deaths may be a cause of their decline in Romania.
One of the recently observed habitats, the
Romanian Plain, is known to be heavily affected by agriculture.
Habitat loss
Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
to agriculture may also be a cause of the species' decline in Romania.
References
{{Taxonbar, from=Q41694
Dolichophis
Reptiles described in 1789
Taxa named by Johann Friedrich Gmelin
Snakes of Jordan
Reptiles of Europe
Reptiles of Russia