Carrion Crow
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The carrion crow (''Corvus corone'') is a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird of the family
Corvidae Corvidae is a Cosmopolitan distribution, cosmopolitan Family (biology), family of Songbird, oscine passerine birds that contains the crows, ravens, Rook (bird), rooks, magpies, jackdaws, jays, treepies, choughs, and Nutcracker (bird), nutcrackers ...
, native to western
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and the eastern
Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...
.


Taxonomy and systematics

The carrion crow was one of the many species originally described by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in his landmark 1758 10th edition of ''Systema Naturae'', and it still bears its original name of ''Corvus corone''. The binomial name is derived from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
, "raven", and Greek , "crow". The hooded crow (''Corvus cornix''), formerly regarded as a subspecies, has been split off as a separate species, and there is some discussion whether the eastern carrion crow (''C. c. orientalis'') is distinct enough to warrant specific status; the two taxa are well separated, and it has been proposed they could have evolved independently in the wetter, maritime regions at the opposite ends of the
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
n landmass.Madge, Steve & Burn, Hilary (1994): ''Crows and jays: a guide to the crows, jays and magpies of the world''. A&C Black, London. Along with the hooded crow, the carrion crow occupies a similar ecological niche in Eurasia to the American crow (''Corvus brachyrhynchos'') in North America. The two species look very similar to one another, but can be differentiated by size, as the carrion crow is larger and of a stockier build compared to the American crow.


Description

The plumage of the carrion crow is black with a green or purple sheen, much greener than the gloss of the rook (''Corvus frugilegus''). The bill, legs and feet are also black. It can be distinguished from the
common raven The common raven or northern raven (''Corvus corax'') is a large all-black passerine bird. It is the most widely distributed of all Corvidae, corvids, found across the Northern Hemisphere. There are 11 accepted subspecies with little variatio ...
by its size of around in length as compared to an average of for ravens, and from the hooded crow by its black plumage. The carrion crow has a wingspan of and weighs . Juvenile carrion crows can be identified by their brownish plumage and blue eyes, both of which darken to black and brown as the crow grows older.


Distribution and genetic relationship to hooded crows

The carrion crow (''Corvus corone'') and hooded crow (''Corvus cornix''), including the former's slightly larger allied
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
''C. c. orientalis'', are two very closely related species; the geographic distributions of both forms of carrion crow across Europe are illustrated in the accompanying diagram. It is believed that this distribution might have resulted from the glaciation cycles during the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
, which caused the parent population to split into isolates which subsequently re-expanded their ranges when the climate warmed causing secondary contact. Further reading

/ref> Poelstra and coworkers sequenced almost the entire genomes of both species in populations at varying distances from the contact zone to find that the two species were genetically identical, both in their DNA and in its expression (in the form of ''m''RNA), except for the ''lack of expression'' of a small portion (<0.28%) of the genome (situated on avian chromosome 18) in the hooded crow, which imparts the lighter plumage colouration on its torso. Thus the two species can viably hybridize, and occasionally do so at the contact zone, but the all-black carrion crows on the one side of the contact zone mate almost exclusively with other all-black carrion crows, while the same occurs among the hooded crows on the other side of the contact zone. It is therefore clear that it is only the outward appearance of the two species that inhibits hybridization. The authors attribute this to assortative mating (rather than to ecological selection), the advantage of which is not clear, and it would lead to the rapid appearance of streams of new lineages, and possibly even species, through mutual attraction between mutants. Unnikrishnan and Akhila propose, instead, that koinophilia is a more parsimonious explanation for the resistance to hybridization across the contact zone, despite the absence of physiological, anatomical or genetic barriers to such hybridization. The carrion crow is also found in the mountains and forests of Japan and also in the cities of Japan.


Behaviour and ecology

The rook is generally gregarious and the crow largely solitary, but rooks occasionally nest in isolated trees, and crows may feed with rooks; moreover, crows are often sociable in winter roosts. The most distinctive feature is the voice. The rook has a high-pitched ''kaaa'', but the crow's guttural, slightly vibrant, deeper croaked ''kraa'' is distinct from any note of the rook. The carrion crow is noisy, perching on a vantage point such as a building or the top of a tree and calling three or four times in quick succession, with a slight pause between each series of croaks. During each series of calls, a crow may perform an accompanying gesture, raising its shoulders and bowing its head and neck downwards with each caw. The wing-beats are slower, more deliberate than those of the rook. Carrion crows can become tame near humans, and can often be found near areas of human activity or habitation including cities, moors, woodland, sea cliffs and farmland where they compete with other social birds such as
gull Gulls, or colloquially seagulls, are seabirds of the subfamily Larinae. They are most closely related to terns and skimmers, distantly related to auks, and even more distantly related to waders. Until the 21st century, most gulls were placed ...
s, other corvids, and
duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
s for food in parks and gardens. File:Bois de Vincennes, Carrion crow cawing and bowing.webm, The carrion crow can be seen performing bowing gestures while cawing. File:Bois de Vincennes, Carrion crows vocalizing.webm, Other vocalizations of the carrion crow include clicking. Like other species of corvid, carrion crows will actively harass predators and competitors that enter their territory or threaten them or their offspring, and will engage in group mobbing behaviour as a method to defend themselves.


Intelligence

Like all corvids, carrion crows are very intelligent. For example, they can discriminate between numerosities up to 30, flexibly switch between rules, and recognise human and crow faces. Given the difference in brain architecture in crows compared to primates, these abilities suggest that their intelligence is realised as a product of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
.


Diet

Though an eater of
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
of all kinds, the carrion crow will eat
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s,
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s, other
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s, grain, fruits, seeds, nuts, small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s,
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s,
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
, scraps and will also steal eggs. Crows are scavengers by nature, which is why they tend to frequent sites inhabited by
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s in order to feed on their household waste. Crows will also harass
birds of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
or even foxes for their kills. Crows actively hunt and occasionally co-operate with other crows to make kills, and are sometimes seen catching
duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
lings for food. Due to their gregarious lifestyle and defensive abilities, carrion crows have few natural predators. However, powerful raptors such as the Eurasian goshawk,
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known simply as the peregrine, is a Cosmopolitan distribution, cosmopolitan bird of prey (raptor) in the family (biology), family Falconidae renowned for its speed. A large, Corvus (genus), cro ...
,
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl, a type of bird that resides in much of Eurasia. It is often just called the eagle-owl in Europe and Asia. It is one of the largest species of owl. Females can grow to a total le ...
and
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of pr ...
will readily hunt them, and crows can become an important prey item locally.


Nesting

The bulky stick nest is usually placed in a tall tree, but cliff ledges, old buildings and pylons may be used as well. Nests are also occasionally placed on or near the ground. The nest resembles that of the
common raven The common raven or northern raven (''Corvus corax'') is a large all-black passerine bird. It is the most widely distributed of all Corvidae, corvids, found across the Northern Hemisphere. There are 11 accepted subspecies with little variatio ...
, but is less bulky. The 3 to 4 brown-speckled blue or greenish eggs are incubated for 18–20 days by the female alone, who is fed by the male. The young fledge after 29–30 days.British Trust for Ornithology (2005) Nest Record Scheme data
It is not uncommon for an offspring from the previous years to stay around and help rear the new hatchlings. Instead of seeking out a mate, it looks for food and assists the parents in feeding the young.


References


External links

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Photo of profile

Image of the skull

Ageing and sexing (PDF; 3.4 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze

HOME of the corvus corone corone
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