Carphephorus Carnosus
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''Carphephorus'' is a genus of
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
n plants in the family
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
. They are native to the southeastern
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
from
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
to
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
. Plants of this genus are known commonly as chaffheads.''Carpephorus''.
Integrated Taxonomic Information System (ITIS).


Description

These are
perennial plant In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...
s that grow from a
caudex A caudex (: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695 In the strict sense of the term, meaning a stem, "caudex" is most ...
and fibrous root system. The stems are erect and unbranched, usually reaching 20 to 60 centimeters (8-24 inches) in height, and taller at times. The leaves are
alternate Alternative or alternate may refer to: Arts, entertainment and media * Alternative (''Kamen Rider''), a character in the Japanese TV series ''Kamen Rider Ryuki'' * Alternative comics, or independent comics are an alternative to mainstream superh ...
ly arranged and point upward, sometimes pressed against the stem. The blades vary in shape and are hairy to hairless and generally
glandular A gland is a cell or an organ in an animal's body that produces and secretes different substances that the organism needs, either into the bloodstream or into a body cavity or outer surface. A gland may also function to remove unwanted substance ...
. The
flower heads A pseudanthium (; : pseudanthia) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, composite flowers ...
are borne in open
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s. Each head contains up to about 35
disc floret Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger fa ...
s, usually
lavender ''Lavandula'' (common name lavender) is a genus of 47 known species of perennial flowering plants in the sage family, Lamiaceae. It is native plant, native to the Old World, primarily found across the drier, warmer regions of the Mediterranean ...
to dark
magenta Magenta () is a purple-red color. On color wheels of the RGB color model, RGB (additive) and subtractive color, CMY (subtractive) color models, it is located precisely midway between blue and red. It is one of the four colors of ink used in colo ...
or pinkish purple, sometimes blue. The fruit is a ribbed, rough-textured cypsela with a pappus of bristles.


Classification

Some authors separate certain species into separate genera, ''Trilisa'' and ''Litrisa'', on the basis of certain floral characters. The species are similar enough in other aspects that other authors maintain them in ''Carphephorus''. Molecular data may support the separation of at least some of the taxa. ''Carphephorus'' is in the tribe
Eupatorieae Eupatorieae is a Tribe (biology), tribe of over 2000D.J.N.Hind & H.E.Robinson. 2007. Tribe Eupatorieae In: ''The Families and Genera of Vascular Plants'' vol.VIII. (Joachim W.Kadereit & Charles Jeffrey, volume editors. Klaus Kubitzky, general edi ...
of the
aster family Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger fam ...
. Like other members of this tribe, the
flower heads A pseudanthium (; : pseudanthia) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, composite flowers ...
have
disc floret Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger fa ...
s and no
ray florets Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger fa ...
. It is also in the subtribe Liatrinae along with, for example, ''
Liatris ''Liatris'' (), commonly known as gayfeather and blazing star is a genus of flowering plants in the tribe Eupatorieae within the family Asteraceae native to North America (Canada, United States, Mexico and the Bahamas). Some species are used ...
'' and ''
Garberia ''Garberia'' is a monotypic taxon, monotypic genus of flowering plants in the family Asteraceae, containing the single species ''Garberia heterophylla''. ;Species and varietiesFlann, C (ed) 2009+ Global Compositae Checklist
/ref> *'' Carphephorus bellidifolius'' – sandywoods chaffhead -
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
*'' Carphephorus carnosus'' (syn. ''Trilisa carnosa'') – pineland chaffhead -
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
*'' Carphephorus corymbosus'' – coastal plain chaffhead -
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
*''
Carphephorus odoratissimus ''Carphephorus odoratissimus'' (syn. ''Trilisa odoratissima''), common name vanillaleaf, is a species of North American plants in the family Asteraceae. This species is native to the southeastern United States, including the states of Georgia, No ...
'' (syn. ''Trilisa odoratissima'') – vanillaleaf -
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
**''C. odoratissimus'' var. ''odoratissimus'' **''C. odoratissimus'' var. ''subtropicanus'' (sometimes treated as a separate species, ''Carphephorus subtropicanus''). This species or variety is more southern than ''C. odoratissimus'' var. ''odoratissimus'', lacks the characteristic
coumarin Coumarin () or 2''H''-chromen-2-one is an aromatic organic chemical compound with formula . Its molecule can be described as a benzene molecule with two adjacent hydrogen atoms replaced by an unsaturated lactone ring , forming a second six-me ...
odor of the latter, and has a different growth habit, being smaller with more of a
rosette Rosette is the French diminutive of ''rose''. It may refer to: Flower shaped designs * Rosette (award), a mark awarded by an organisation * Rosette (design), a small flower design *hence, various flower-shaped or rotational symmetric forms: ** R ...
form. There is some overlap between the ranges of the two. *'' Carphephorus pseudoliatris'' – bristleleaf chaffhead -
Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
*'' Carphephorus tomentosus'' – woolly chaffhead -
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
;Formerly included *'' Carphephorus paniculatus'' (syn of '' Trilisa paniculata '') – hairy chaffhead -
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...


Biochemistry

A number of species contain
volatile oils An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
, giving characteristic odors.


References

{{Taxonbar, from=Q5045796 Eupatorieae Asteraceae genera Flora of the Southeastern United States Taxa named by Henri Cassini