''Caroxylon imbricatum'',

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

''Salsola imbricata'', is a small

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of shrub in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Amaranthaceae Amaranthaceae ( ) is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, maki ...

. It grows in deserts and arid regions of north Africa, the Arabian Peninsula and southwestern Asia.

Description

''Caroxylon imbricatum'' is a small, spreading shrub or sub-shrub growing up to tall. The grey or reddish stems are up to thick and these and the lower leaves are densely hairy. In the upper parts of the plant the stems are creamy or pale grey and branch frequently, some branches growing vertically while others spread horizontally. Regularly-arranged, catkin-like branchlets project from the branches. The leaves are tiny, succulent and linear or narrowly triangular. The inflorescence is spike-like with bracts similar to the leaves, small flowers with five petals, five stamens and two styles. The fruiting perianth has silky wings.

Taxonomy

The species was first described in 1775 as ''Salsola imbricata'' by the Swedish naturalist

Peter Forsskål Peter Forsskål, sometimes spelled Pehr Forsskål, Peter Forskaol, Petrus Forskål or Pehr Forsskåhl (11 January 1732 – 11 July 1763) was a Sweden, Swedish exploration, explorer, oriental studies, orientalist, natural history, naturalist, and ...

. In 1849,

Alfred Moquin-Tandon Christian Horace Benedict Alfred Moquin-Tandon (7 May 1804 – 15 April 1863) was a French Natural history, naturalist and doctor. Moquin-Tandon was professor of zoology at Marseille from 1829 until 1833, when he was appointed professor of botany ...

transferred it to the genus ''

Caroxylon ''Caroxylon'' is a genus of shrubby flowering plants in the family Amaranthaceae, found in drier areas of the Old World, including southern Africa, Madagascar, northern Africa, Mediterranean islands of Europe, the Canary Islands, Socotra, Ukraine ...

'', making it ''Caroxylon imbricatum'' (Forssk.) Moq., but later it was mostly accepted in genus ''Salsola''. Following a phylogenetic analysis of Salsoloideae in 2007 by Akhani, H., et al., it has been proposed to place ''Salsola imbricata'' back into ''Caroxylon imbricatum''. This placement is accepted by GBIF and

Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...

Distribution and habitat

This plant has a widespread distribution across the desert belt of Saharan Africa, the Arabian Peninsula, southern Iran, Pakistan, Afghanistan and northwestern India. It typically grows in disturbed areas such as runnels, washes, dry wadis, eroded slopes and coastal cliffs. It grows on various soil types and is a

ruderal species A ruderal species is a plant species that is first to colonize disturbed lands. The disturbance may be natural for example, wildfires or avalanchesor the consequences of human activities, such as construction ( of roads, of buildings, mining, e ...

, colonising fallow land and over-grazed pastures.

Ecology

''Caroxylon imbricatum'' is a

halophytic A halophyte is a salt-tolerant plant that grows in soil or waters of high salinity, coming into contact with saline water through its roots or by salt spray, such as in saline semi-deserts, mangrove swamps, marshes and sloughs, and seashores. ...

plant; under conditions of salt stress, the plant increases its water content (becomes more succulent) and decreases the surface area of its leaves. Tests on the germination rates of seeds show that ''Caroxylon imbricatum'' sprouts more quickly and consistently at 20 °C than at higher temperatures, and shows higher germination rates at lower salinity levels than high ones. However, seeds treated at high salinity levels recovered their germination potential after immersion in unsalted water. The species has traditionally been used as a

vermifuge Anthelmintics or antihelminthics are a group of antiparasitic drugs that expel parasitic worms (helminths) and other internal parasites from the body by either stunning or killing them without causing significant damage to the host. They may also ...

and for treating certain skin disorders. Five

triterpene Triterpenes are a class of terpenes composed of six isoprene units with the molecular formula C30H48; they may also be thought of as consisting of three terpene units. Animals, plants and fungi all produce triterpenes, including squalene, the pre ...

glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...

s have been isolated from the roots of ''Caroxylon imbricatum'', two of them being new glycoside derivatives not previously known.

References

{{Taxonbar, from=Q17246081, from2=Q12213843 Amaranthaceae Desert flora Flora of the Arabian Peninsula Flora of North Africa