The Carolinian forest refers to a life zone in eastern North America characterized primarily by the predominance of

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...

(broad-leaf) forest. The term "Carolinian", which is most commonly used in Canada, refers to the deciduous forests which span across much of the eastern United States from the

Carolinas The Carolinas are the U.S. states of North Carolina and South Carolina, considered collectively. They are bordered by Virginia to the north, Tennessee to the west, and Georgia to the southwest. The Atlantic Ocean is to the east. Combining Nort ...

northward into

southern Ontario Southern Ontario is a primary region of the province of Ontario, Canada, the other primary region being Northern Ontario. It is the most densely populated and southernmost region in Canada. The exact northern boundary of Southern Ontario is dis ...

, Canada. These deciduous forests in the United States and southern Ontario share many similar characteristics and species hence their association. Today the term is often used to refer to the Canadian portion (northern limit) of the deciduous forest region while the portion in the United States is often referred to as the "Eastern deciduous forest".

Location and extent

The Carolinian zone spans across much of the eastern United States, with extensive coverage in the

, the

Virginias The Virginias (sometimes also known as the two Virginias) is a region in the United States comprising the U.S. states of Virginia and West Virginia. If they were a single state (as they were until the Civil War), the Virginias would have a c ...

Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virgini ...

Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to ...

Maryland Maryland ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; ...

Delaware Delaware ( ) is a state in the Mid-Atlantic region of the United States, bordering Maryland to its south and west; Pennsylvania to its north; and New Jersey and the Atlantic Ocean to its east. The state takes its name from the adjacen ...

Pennsylvania Pennsylvania (; (Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, Ma ...

, parts of southern

New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...

state,

Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York to the west, and Long Island Sound to the south. Its cap ...

, and

Rhode Island Rhode Island (, like ''road'') is a state in the New England region of the Northeastern United States. It is the smallest U.S. state by area and the seventh-least populous, with slightly fewer than 1.1 million residents as of 2020, but i ...

, eastern

Ohio Ohio () is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Of the List of states and territories of the United States, fifty U.S. states, it is the List of U.S. states and territories by area, 34th-l ...

, and small parts of southern

Michigan Michigan () is a U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the List of U.S. states and ...

Indiana Indiana () is a U.S. state in the Midwestern United States. It is the 38th-largest by area and the 17th-most populous of the 50 States. Its capital and largest city is Indianapolis. Indiana was admitted to the United States as the 19th ...

, and western

. It extends up into

Southern Ontario Southern Ontario is a primary region of the province of Ontario, Canada, the other primary region being Northern Ontario. It is the most densely populated and southernmost region in Canada. The exact northern boundary of Southern Ontario is dis ...

, Canada which is located in the fertile ecozone of the Mixedwood Plains and includes ecodistricts 7E-1 to 7E-6. Trees found here include various species of

ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...

chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelate ...

hickory Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in M ...

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

, and

walnut A walnut is the edible seed of a drupe of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. Although culinarily considered a "nut" and used as such, it is not a tru ...

; tallest of all is the

tulip tree ''Liriodendron'' () is a genus of two species of characteristically large trees, deciduous over most of their populations, in the magnolia family ( Magnoliaceae). These trees are widely known by the common name tulip tree or tuliptree for their ...

. Fruit trees native to this zone include the pawpaw. Animal life includes

raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight o ...

possums Possum may refer to: Animals * Phalangeriformes, or possums, any of a number of arboreal marsupial species native to Australia, New Guinea, and Sulawesi ** Common brushtail possum (''Trichosurus vulpecula''), a common possum in Australian urban ...

, squirrels (including the relatively rare

southern flying squirrel The southern flying squirrel or the assapan (''Glaucomys volans'') is one of three species of the genus '' Glaucomys'' and one of three flying squirrel species found in North America. It is found in deciduous and mixed woods in the eastern half ...

nuthatch The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. Mo ...

es, and

chickadee The chickadees are a group of North American birds in the tit family included in the genus '' Poecile''. Species found in North America are referred to as chickadees, while other species in the genus are called tits. They are small-sized bird ...

Carolinian Canada

The Carolinian forest in Canada is located at the southern tip of Ontario between Lake Erie, Lake Huron and Lake Ontario. The region contains an extremely high

biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity' ...

of species, over 500 of which are considered rare.

Climate

The reason for the high biodiversity in this region is its unique climate; the Carolinian forest of Ontario has the warmest average annual temperatures, the longest frost-free seasons, and the mildest winters of any region in Ontario. This distinctive climate is largely due to the nearby

Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five la ...

which moderate the temperature of the surrounding land.

Status

The high

fertility Fertility is the capability to produce offspring through reproduction following the onset of sexual maturity. The fertility rate is the average number of children born by a female during her lifetime and is quantified demographically. Ferti ...

of the land has seen the region become highly developed and populated, with agricultural, industrial, commercial and urban areas. Today, the Carolinian Zone contains major cities and is home to one quarter of Canada's population despite being 0.25% of the total land area. The deforestation of the region for this development has led to significant

habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

and

fragmentation Fragmentation or fragmented may refer to: Computers * Fragmentation (computing), a phenomenon of computer storage * File system fragmentation, the tendency of a file system to lay out the contents of files non-continuously * Fragmented distributi ...

, leaving the remaining portions of land scattered and disconnected, with some areas still threatened by human development. In total, it is estimated that forest cover has been reduced from 80% to 11.3% while wetlands reduced from 28.3% to 5.1%. In addition to habitat loss and fragmentation, the native Carolinian species are also being threatened by invasive species such as

garlic mustard ''Alliaria petiolata'', or garlic mustard, is a Biennial plant, biennial flowering plant in the mustard family (Brassicaceae). It is native to Europe, western and central Asia, north-western Africa, Morocco, Iberian Peninsula, Iberia and the ...

and buckthorn, and

overgrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature ...

White-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...

. A list of invasive species in the Carolinian zone can be foun
here
These factors have contributed to the Carolinian zone becoming the most threatened region in Ontario, with over 125 species of plants and animals listed as either vulnerable, threatened or endangered by the federal or provincial governments. This is over one-third of all

vulnerable Vulnerable may refer to: General * Vulnerability * Vulnerability (computing) * Vulnerable adult * Vulnerable species Music Albums * ''Vulnerable'' (Marvin Gaye album), 1997 * ''Vulnerable'' (Tricky album), 2003 * ''Vulnerable'' (The Used album) ...

threatened Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensa ...

endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...

species in Canada. A full list of these species can be foun
here

Conservation efforts

Some parts of the remaining natural area in the Carolinian zone are protected in an effort to conserve the region and its unique, diverse biota. For example, today there are many protected areas including Point Pelee National Park, 21 provincial parks, and many conservation areas. Some of the best preserved areas of Canada's Carolinian forest are located in Windsor's

Ojibway Prairie Complex The Ojibway Prairie Complex is a 350- hectare complex of parks and nature reserves on the west side of Windsor, Ontario, Canada. It comprises Ojibway Park, Tallgrass Prairie Heritage Park, Black Oak Heritage Park, and the Spring Garden Natur ...

and Rondeau Provincial Park near Morpeth, Ontario; another is the

Niagara Glen Nature Reserve Niagara Glen Nature Reserve is a nature reserve located near the Niagara Whirlpool along the Canadian side of the Niagara Gorge. It is one of the best preserved remaining example of Southern Ontario's original Carolinian forest. The park is oper ...

near

Niagara Falls, Ontario Niagara Falls is a city in Ontario, Canada. It is on the western bank of the Niagara River in the Golden Horseshoe region of Southern Ontario, with a population of 88,071 at the Canada 2016 Census, 2016 census. It is part of the List of census ...

. There are also organizations like th
Carolinian Canada Coalition
who aim to restore the region as much as possible. Carolinian forests are complex ecosystems that cannot be replicated in gardens or other small areas. The only way to preserve this unique habitat is continue to keep larger areas of land protected from development and agriculture. Other conservation efforts have included the reduction of the white-tailed deer population from their peak density of 55 deer km⁻² to 7 deer km^−2, between the years of 1996-2009. However, studies have found that while this helps reduce further forest damage, continued decline in the forest canopy can still occur, indicating the recovery from overgrazing by herbivores on the forest canopy is a long process without immediate results. Maintaining a lower density of white-tailed deer, while increasing the seed sources of native trees and protecting tree saplings in herbivore exclosures are all suggested ways of helping the Carolinian forest recover or at least conserve what is remaining.

Examples of species

Fauna: *

American black bear The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bea ...

Virginia opossum The Virginia opossum (''Didelphis virginiana''), also known as the North American opossum, is the only opossum living north of Mexico, its range extending south into Central America. It is the northernmost marsupial in the world. In the United S ...

American badger The American badger (''Taxidea taxus'') is a North American badger similar in appearance to the European badger, although not closely related. It is found in the western, central, and northeastern United States, northern Mexico, and south-cen ...

Gray fox The gray fox (''Urocyon cinereoargenteus''), or grey fox, is an omnivorous mammal of the family Canidae, widespread throughout North America and Central America. This species and its only congener (biology), congener, the diminutive island fox ...

Bobcat The bobcat (''Lynx rufus''), also known as the red lynx, is a medium-sized cat native to North America. It ranges from southern Canada through most of the contiguous United States to Oaxaca in Mexico. It is listed as Least Concern on the IU ...

Southern flying squirrel The southern flying squirrel or the assapan (''Glaucomys volans'') is one of three species of the genus '' Glaucomys'' and one of three flying squirrel species found in North America. It is found in deciduous and mixed woods in the eastern half ...

Wild turkey The wild turkey (''Meleagris gallopavo'') is an upland ground bird native to North America, one of two extant species of turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic turkey, which was originally ...

Ruffed grouse The ruffed grouse (''Bonasa umbellus'') is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is the most widely distributed game bird in North America. It is non-migratory. It is the only specie ...

* Hooded warbler *

Prothonotary warbler The prothonotary warbler (''Protonotaria citrea'') is a small songbird of the New World warbler family. It is named for its plumage which resembles the yellow robes once worn by papal clerks (named prothonotaries) in the Roman Catholic church. ...

* Canada Warbler *

Carolina wren The Carolina wren (''Thryothorus ludovicianus'') is a common species of wren that is a resident in the eastern half of the United States of America, the extreme south of Ontario, Canada, and the extreme northeast of Mexico. Severe winters restri ...

Yellow-breasted chat The yellow-breasted chat (''Icteria virens'') is a large songbird found in North America, and is the only member of the family Icteriidae. It was once a member of the New World warbler family, but in 2017, the American Ornithological Society move ...

Northern cardinal The northern cardinal (''Cardinalis cardinalis'') is a bird in the genus '' Cardinalis''; it is also known colloquially as the redbird, common cardinal, red cardinal, or just cardinal (which was its name prior to 1985). It can be found in southe ...

* Red bellied woodpecker *

Red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members wit ...

Barn owl The barn owl (''Tyto alba'') is the most widely distributed species of owl in the world and one of the most widespread of all species of birds, being found almost everywhere except for the polar and desert regions, Asia north of the Himala ...

Eastern box turtle The eastern box turtle (''Terrapene carolina carolina'') is a subspecies within a group of hinge-shelled turtles normally called box turtles. ''T. c. carolina'' is native to the eastern part of the United States. The eastern box turtle ...

Spotted turtle The spotted turtle (''Clemmys guttata''), the only species of the genus '' Clemmys'', is a small, semi-aquatic turtle that reaches a carapace length of upon adulthood. Their broad, smooth, low dark-colored upper shell, or carapace, ranges in it ...

Eastern hognose snake The eastern hog-nosed snake (''Heterodon platirhinos''), also known as the spreading adder Wright, A.H., and A.A. Wright (1957). ''Handbook of Snakes of the United States and Canada''. (in 2 volumes). Ithaca and London: Comstock Publishing Assoc ...

*Eastern spiny softshell turtle *

Pantherophis gloydi ''Pantherophis vulpinus'', commonly known as the eastern foxsnake or eastern fox snake, is a species of rat snake in the family Colubridae. The species is nonvenomous and is endemic to the eastern Great Lakes region of the United States, as well ...

* Red salamander * Karner blue butterfly Flora: * Eastern prickly pear cactus *

Tulip tree ''Liriodendron'' () is a genus of two species of characteristically large trees, deciduous over most of their populations, in the magnolia family ( Magnoliaceae). These trees are widely known by the common name tulip tree or tuliptree for their ...

Sassafras ''Sassafras'' is a genus of three extant and one extinct species of deciduous trees in the family Lauraceae, native to eastern North America and eastern Asia.Wolfe, Jack A. & Wehr, Wesley C. 1987. The sassafras is an ornamental tree. "Middle E ...

* Miami mist *

Kentucky Coffee Tree The Kentucky coffeetree (''Gymnocladus dioicus''), also known as American coffee berry, Kentucky mahogany, nicker tree, and stump tree, is a tree in the subfamily Caesalpinioideae of the legume family Fabaceae, native to the Midwest, Upper Sout ...

Flowering Dogwood ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florid ...

Black Walnut ''Juglans nigra'', the eastern American black walnut, is a species of deciduous tree in the walnut family, Juglandaceae, native to North America. It grows mostly in riparian zones, from southern Ontario, west to southeast South Dakota, ...

American Chestnut The American chestnut (''Castanea dentata'') is a large, fast-growing deciduous tree of the beech family native to eastern North America. As is true of all species in genus Castanea, the American chestnut produces burred fruit with edible nuts. ...

*Sycamore *

Red Mulberry ''Morus rubra'', commonly known as the red mulberry, is a species of mulberry native to eastern and central North America. It is found from Ontario, Minnesota, and Vermont south to southern Florida, and west as far as southeastern South Dak ...

* Black Gum *

Red Maple ''Acer rubrum'', the red maple, also known as swamp maple, water maple, or soft maple, is one of the most common and widespread deciduous trees of eastern and central North America. The U.S. Forest Service recognizes it as the most abundant nati ...

Eastern White Pine ''Pinus strobus'', commonly called the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland (island), Newfoundland, Canada west ...

Eastern Hemlock ''Tsuga canadensis'', also known as eastern hemlock, eastern hemlock-spruce, or Canadian hemlock, and in the French-speaking regions of Canada as ''pruche du Canada'', is a coniferous tree native to eastern North America. It is the state tree ...

Rivers and creeks

Ausable River Au Sable or Ausable may refer to various places: Michigan * Au Sable Township, Iosco County, Michigan ** Au Sable, Michigan, an unincorporated community in the above township * Au Sable Township, Roscommon County, Michigan New York *Au Sable, Ne ...

* Catfish Creek *

Credit River The Credit River is a river in southern Ontario, which flows from headwaters above the Niagara Escarpment near Orangeville and Caledon East to empty into Lake Ontario at Port Credit, Mississauga. It drains an area of approximately . The total ...

* Kettle Creek * Grand River * Rouge River *

Sydenham River The Sydenham River is a river in Chatham-Kent, Lambton County and Middlesex County in southwestern Ontario, Canada, flowing southwest from its source west of London, Ontario and emptying into Lake Saint Clair. The length of the river is and it ...

Thames River The River Thames ( ), known alternatively in parts as the River Isis, is a river that flows through southern England including London. At , it is the longest river entirely in England and the second-longest in the United Kingdom, after the ...

* Twenty Mile Creek *

Welland River The Welland River (originally called the Chippawa Creek) is a river in the Golden Horseshoe that passes through the Southern Ontario cities of Welland and Niagara Falls. It flows from its source just south of Hamilton, Ontario to meet the Nia ...

Nith River The Nith River is a river in Brant, Oxford and Perth Counties and the Regional Municipality of Waterloo in Southwestern Ontario, Canada. The Nith River is approximately 125km in length and empties into the Grand River at the town of Paris. ...

Other

Rouge Park Rouge National Urban Park is a national urban park in Ontario, Canada. The park is centred around the Rouge River and its tributaries in the Greater Toronto Area. The southern portion of the park is situated around the mouth of river in ...

* Barker's Bush *

Point Pelee National Park Point Pelee National Park (; french: Parc national de la Pointe-Pelée) is a national park in Essex County in southwestern Ontario, Canada where it extends into Lake Erie. The word is French for 'bald'. Point Pelee consists of a peninsula of la ...

References

{{Reflist

External links

Carolinian Canada
Temperate broadleaf and mixed forests in the United States Ecoregions of the United States Ecoregions of Canada Forests of Ontario Forests of the United States

Flora of the Eastern United States Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring ( indigenous) native plants. The corresponding term for animals is ''fauna'', and for fungi, it is '' funga''. Sometimes ...

Temperate broadleaf and mixed forests in Canada