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Carnivoramorpha ("carnivoran-like forms") is a clade of placental mammals that includes the modern order
Carnivora Carnivora is a monophyletic order of placental mammals consisting of the most recent common ancestor of all cat-like and dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, ...
and its extinct stem-relatives.Bryant, H.N., and M. Wolson (2004
“Phylogenetic Nomenclature of Carnivoran Mammals.”
''First International Phylogenetic Nomenclature Meeting''. Paris, Museum National d’Histoire Naturelle


Classification and phylogeny


Traditional classification

* Clade: Carnivoramorpha
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
clades to Carnivora, but without Creodonts] ** Order:
Carnivora Carnivora is a monophyletic order of placental mammals consisting of the most recent common ancestor of all cat-like and dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, ...
(carnivorans) *** Suborder:
Caniformia Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs ( wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia ( seals, walruses and sea lions) are also assigned to this gro ...
("dog-like" carnivorans) *** Suborder:
Feliformia Feliformia is a suborder within the order Carnivora consisting of "cat-like" carnivorans, including cats (large and small), hyenas, mongooses, viverrids, and related taxa. Feliformia stands in contrast to the other suborder of Carnivora, Canifor ...
("cat-like" carnivorans) *** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': **** Genus: †''
Palaeogale ''Palaeogale'' is an extinct genus of carnivorous mammal known from the Late Eocene, Oligocene, and Early Miocene of North America, Europe, and Eastern Asia. A small carnivore often associated with the mustelids, ''Palaeogale'' might have been s ...
'' ** Superfamily: † Miacoidea *** Family: † Miacidae *** Family: † Viverravidae *** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': **** †'' "Sinopa" insectivorus'' ** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': *** Genus: †'' Ravenictis'' *** † Carnivoramorpha sp. (''UALVP 31176'')


Revised classification

Recent phylogenetic studies indicate that the superfamily Miacoidea and family Miacidae are
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
, with "miacids" being more closely related to carnivorans than to viverravids. In 2010 Flynn, Finarelli & Spaulding named a new clade
Carnivoraformes Carnivoraformes ("carnivoran-like forms") is a clade of placental mammals that includes the modern order Carnivora and its extinct stem-relatives. Classification and phylogeny Classification In 2010 Flynn, Finarelli & Spaulding named a new clade ...
within Carnivoramorpha, containing carnivorans and "miacids" but not viverravids. The authors defined Carnivoraformes as the clade containing Carnivora and all taxa that are more closely related to Carnivora (represented by ''
Canis lupus The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
'') than to viverravids (represented by '' Viverravus gracilis''). * ''Clade'': Carnivoramorpha ** ''Clade'':
Carnivoraformes Carnivoraformes ("carnivoran-like forms") is a clade of placental mammals that includes the modern order Carnivora and its extinct stem-relatives. Classification and phylogeny Classification In 2010 Flynn, Finarelli & Spaulding named a new clade ...
*** Order:
Carnivora Carnivora is a monophyletic order of placental mammals consisting of the most recent common ancestor of all cat-like and dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, ...
(carnivorans) *** Family: † Quercygalidae *** (unranked): †''Gracilocyon''/''Oodectes'' clade **** Genus: †'' Eogale'' **** Genus: †''
Gracilocyon ''Gracilocyon'' is an extinct genus of miacid carnivoran which existed in United States, Belgium and in the United Kingdom during the earliest Eocene. It was first named by Thierry Smith and Richard Smith in 2010 and the type species is ''Graci ...
''(
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
genus)
**** Genus: †''
Oodectes ''Oodectes'' is an extinct genus of Miacidae. Notes Sourcesfindarticles.com*''Fossil Mammalia of the Huerfano Formation, Eocene, of Colorado'' by Peter Robinsonnmnaturalhistory.orgParamiacis ''Paramiacis'' is an extinct genus of Miacidae. Christian Mathis has defined to make a difference between the miacids from Europe and the American genus ''Miacis''. There are two species of it, ''P. exilis'' (Henri Filhol, 1876) and ''P. teilhar ...
'' **** Genus: †''
Paroodectes ''Paroodectes'' is a miacid animal that lived during the early Eocene (ca. 50 million years ago) in the rain forests and swamps of the present-day Germany. It was a prehistoric predator that had the size and the appearance of a cat and was well a ...
'' **** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': ***** † ''"Miacis"'' sp. 'CM 67873'' & ''CM 77299'' *** (unranked): †''Vulpavus'' clade **** Genus: †''
Palaearctonyx ''Palaearctonyx'' is an extinct genus of omnivorous Miacidae which inhabited North America during the Eocene living from 50.3 to 46.2 Ma and existed for approximately . Taxonomy ''Palaearctonyx'' was named by Matthew (1909). Its type is ''Pal ...
'' **** Genus: †''
Vassacyon ''Vassacyon'' is an extinct genus of Miacidae. It contains two species; ''Vassacyon promicrodon'' and ''Vassacyon bowni''. It is considered the largest of the early Eocene mammals. Notes References *Flynn, J.J., 1998. Early Cenozoic Carnivor ...
'' **** Genus: †''
Vulpavus ''Vulpavus'' ("fox grandfather") is an extinct genus of Miacidae. It measured 60–90 cm in length and had an estimated weight over 1.19 kg (in ''V. palustris'').http://museumu03.museumwww.naturkundemuseum-berlin.de/cgi-bin/bridge.pl?a=ba ...
''(paraphyletic genus) *** Genus: †'' Africtis'' *** Genus: †'' Ceruttia'' *** Genus: †''
Dawsonicyon ''Dawsonicyon'' ("Dawson's dog") is an extinct genus of placental mammals from clade Carnivoraformes, that lived in North America during middle Eocene. Fossils of type species ''Dawsonicyon isami'' are known from the ‘Bridger B’, site of Bl ...
'' *** Genus: †'' Dormaalocyon'' *** Genus: †'' Harpalodon'' *** Genus: †'' Lycarion'' *** Genus: †'' Messelogale'' *** Genus: †''
Miacis ''Miacis'' is a genus of extinct carnivorous mammals that appeared in the late Paleocene and continued through the Eocene. The genus ''Miacis'' is not monophyletic but a diverse collection of species that belong to the stemgroup within the ...
'' *** Genus: †''
Miocyon ''Miocyon'' is an extinct genus of Miacidae, primitive carnivores which lived during the Paleocene and Eocene Epoch. Notes References * Flynn, J.J., 1998. Early Cenozoic Carnivora ("Miacoidea"). pp. 110–123 ''in'' C.M. Janis, K.M. Sco ...
'' *** Genus: †'' Neovulpavus'' *** Genus: †''
Procynodictis ''Procynodictis'' ("before '' Cynodictis''") is an extinct genus of placental mammals from clade Carnivoraformes, that lived in North America from the early to middle Eocene The Eocene ( ) Epoch is a geological epoch that lasted from abou ...
'' *** Genus: †''
Prodaphaenus ''Prodaphaenus'' is an extinct genus of Miacidae. The genus has at least one known species: ''Prodaphaenus scotti''. References *zipcodezoo.comSimamphicyon'' *** Genus: †'' Tapocyon'' *** Genus: †''
Uintacyon ''Uintacyon'' is an extinct genus of Miacidae. There are at least two species in the genus; ''Uintacyon rudis'' and ''Uintacyon gingerichi'', the latter being recently discovered. Notes References * Sources *Flynn, J.J., 1998. Early Cenozoi ...
''(paraphyletic genus) *** Genus: †'' Walshius'' *** Genus: †''
Xinyuictis Xinyuictis is an extinct genus of Miacidae. It contains a species, ''Xinyuictis tenuis''. It was once suggested that the species was the same as ''Miacis'', but it was eventually decided that they were different. Notes References * Sources ...
'' *** Genus: †'' Zodiocyon'' *** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': **** †'' "Miacis" boqinghensis'' **** †'' "Miacis" deutschi'' **** †'' "Miacis" exiguus'' **** †'' "Miacis" gracilis'' **** †'' "Miacis" hargeri'' **** †'' "Miacis" hookwayi'' **** †'' "Miacis" invictus'' **** †'' "Miacis" latidens'' **** †'' "Miacis" lushiensis'' **** †'' "Miacis" petilus'' **** † Carnivoraformes undet. Genus A 'UCMP 110072'' **** † Carnivoraformes undet. Genus B 'SDSNH 56335'' *** ichnotaxa of Carnivoraformes: **** Ichnogenus: †'' Falcatipes'' ** Superfamily: †
Viverravoidea Viverravidae ("ancestors of viverrids") is an extinct monophyletic family of mammals from extinct superfamily Viverravoidea within the clade Carnivoramorpha, that lived from the early Palaeocene to the late Eocene in North America, Europe and As ...
*** Family: † Viverravidae ** ''
Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'': *** Genus: †'' Ravenictis'' *** †'' "Sinopa" insectivorus'' *** † Carnivoramorpha sp. (''UALVP 31176'') *** † Carnivoramorpha sp. (''UALVP 50993'' & ''UALVP 50994'') *** † Carnivoramorpha sp. (''USNM 538395'')


Phylogenetic tree

The phylogenetic relationships of Carnivoramorpha are shown in the following cladogram:Prevosti, F. J. & Forasiepi, A. M. (2018
"Introduction. Evolution of South American Mammalian Predators During the Cenozoic: Paleobiogeographic and Paleoenvironmental Contingencies"
Springer Geology. Springer, Cham.


References


Further reading

* C. M. Janis, J. A. Baskin, A. Berta, J. J. Flynn, G. F. Gunnell, R. M. Hunt jr., L. D. Martin, and K. Munthe (1998.) "Carnivorous mammals." In C. M. Janis, K. M. Scott, and L. L. Jacobs (''eds.''

Cambridge University Press, Cambridge. * K. D. Rose and J. D. Archibold (2005) ttps://books.google.ba/books?id=DhchVG_rbQ8C&dq=ravenictis&hl=sr&source=gbs_navlinks_s "The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades" Baltimore and London, Johns Hopkins University Press * K. D. Rose and J. D. Archibold (2005
"Womb with a View: the Rise of Placentals."
In: K. D. Rose and J. D. Archibol
"The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades"
Baltimore and London, Johns Hopkins University Press * * Stiles, David P. (2005
investigation of the ''Vulpes'' and ''Urocyon'' phylogenetic classification: Feliformia or Caniformia?”
Fox Phylogeny. Vertebrate Evolution – Fall 2005, The Evergreen State College, Olympia, WA. * * Wesley-Hunt, Gina D. (2005
“The Morphological Diversification of Carnivores in North America.”
Paleobiology. Vol. 31, Issue 1, pp. 35–55. * Benton, Michael J. and Philip C. J. Donoghue (2007
“Paleontological Evidence to Date the Tree of Life.”
Molecular Biology and Evolution, Volume 24, Issue 1, Pages 26–53 * * Susumu Tomiya, Shawn P. Zack, Michelle Spaulding and John J. Flynn (2019.) "Carnivorous mammals from the Middle Eocene Washakie formation, Wyoming, U.S.A., amd their diversity trajectory in a post-warming world", in
"The Society of Vertebrate Paleontology 79th annual meeting"
* {{paleo-mammal-stub Mammal unranked clades Pan-Carnivora