genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
shrimp
A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...
. They are widely found in
tropical
The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
or
subtropical
The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
water in
Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
,
Oceania
Oceania ( , ) is a region, geographical region including Australasia, Melanesia, Micronesia, and Polynesia. Outside of the English-speaking world, Oceania is generally considered a continent, while Mainland Australia is regarded as its co ...
and
Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
omnivorous
An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
scavenger
Scavengers are animals that consume Corpse decomposition, dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a he ...
carapace
A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
sympatric
In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
, requiring care when interpreting
molecular phylogenetic
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
analyses that do not use a large number of specimens.
As of March 2022, the
Integrated Taxonomic Information System
The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagenc ...
lists the genus ''Caridina'' as having 340 species. These include the following species:
*''
Caridina ablepsia
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Guo, Jiang & Zhang, 1992
*''
Caridina acuta
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Liang, Chen & W.-X. Li, 2005
*''
Caridina acutirostris
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Schenkel, 1902
*''
Caridina africana
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Kingsley, 1883
*''
Caridina alba
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. canton ...
Caridina apodosis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina bakoensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. canton ...
Caridina barakoma
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina boehmei
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina brevidactyla
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Roux, 1919
*''
Caridina breviata
''Caridina'' is a genus of freshwater Atyidae, atyid Caridea, shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter feeding, filter-feeders and omnivorous scavengers. They range from 0.9 to 9 ...
'' N. K. Ng & Cai, 2000
*''
Caridina brevicarpalis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina bruneiana
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina bunyonyiensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina butonensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina choiseul
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina clandestina
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina endehensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina flavilineata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina gabonensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina gracilirostris
''Caridina gracilirostris'' is a species of brackishwater shrimp belong to genus ''Caridina'' in the family Atyidae. It has a wide native range extending from Japan and Fiji through Indonesia to Madagascar.De Grave, S., Wowor, D. & Klotz, W. 2013 ...
Caridina grandirostris
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina huananensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina intermedia
''Caridina'' is a genus of freshwater Atyidae, atyid Caridea, shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter feeding, filter-feeders and omnivorous scavengers. They range from 0.9 to 9 ...
Caridina kunnathurensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina lanceifrons
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina laroeha
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Klotz & von Rintelen, 2013
*''
Caridina leclerci
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina leytensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina maeana
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' de Mazancourt et al. 2020
*''
Caridina maeklongensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Macharoenboon, K., Manonai, V., & Jeratthitikul, E., 2024
*'' Caridina mahalona'' Cai, Wowor & Choy, 2009
*'' Caridina malawensis'' Richard & Clark, 2009
*''
Caridina malayensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Cai, Ng & Choy, 2007
*''
Caridina mariae
''Caridina mariae'', commonly known as the tiger shrimp, is a variety of fresh water, freshwater shrimp from Southeast Asia (neighboring counties of Yingde and Conghua, Guangdong Province, Southern China and Hong Kong SAR) that are popular in ho ...
Caridina mengaeoides
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina meridionalis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina multidentata
''Caridina multidentata'' is a species of shrimp in the family Atyidae. It is native to Japan and Taiwan.De Grave, S., Shy, J. & Cai, X. 2013''Caridina multidentata''. The IUCN Red List of Threatened Species 2013. Downloaded on 14 June 2016. Its ...
'' Stimpson, 1860
*''
Caridina nana
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina nudirostris
''Caridina'' is a genus of freshwater Atyidae, atyid Caridea, shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter feeding, filter-feeders and omnivorous scavengers. They range from 0.9 to 9 ...
Caridina papuana
''Caridina'' is a genus of freshwater Atyidae, atyid Caridea, shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter feeding, filter-feeders and omnivorous scavengers. They range from 0.9 to 9 ...
Caridina paratypus
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina parvirostris
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina pedicultrata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina piokerai
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' de Mazancourt et al. 2020
*''
Caridina pisuku
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' de Mazancourt et al. 2020
*''
Caridina plicata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Liang, 2004
*''
Caridina poarae
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina pseudodenticulata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina pseudoserrata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Đăng & Ðỗ, 2007
*''
Caridina qingyuanensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina serrata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina sikipozo
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina sinanensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina solearipes
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina spelunca
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina spongicola
''Caridina spongicola'' is a small species of freshwater shrimp from Sulawesi (Indonesia) that reaches in length. In the wild it strictly lives on an undescribed species of freshwater sponge, making it one of only two known commensal species of ...
Caridina striata
''Caridina striata'' is a freshwater shrimp from Sulawesi. It is endemic to Lake Poso and Lake Towuti. Common names in the aquarium hobby are red line shrimp and red stripe shrimp. It is commonly found on rocky substrates.
Threats
It is currentl ...
'' von Rintelen & Cai, 2009
*''
Caridina subventralis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina tetrazona
''Caridina tetrazona'' is a species of freshwater shrimp native to the Wanshan islands in Guangdong Province, southern China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, ...
Caridina trifasciata
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina tupaia
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' de Mazancourt, Marquet & Keith, 2019
*''
Caridina turipi
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' de Mazancourt et al. 2020
*''
Caridina typus
''Caridina typus'', also known as the Australian Amano shrimp, is a species of amphidromous
Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales r ...
Caridina venusta
''Caridina'' is a genus of freshwater Atyidae, atyid Caridea, shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter feeding, filter-feeders and omnivorous scavengers. They range from 0.9 to 9 ...
Caridina vitiensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina williamsi
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' Cai & Ng, 2000
*''
Caridina woltereckae
''Caridina woltereckae'', or Sulawesi harlequin shrimp as it is commonly known in the aquarium hobby, is a freshwater shrimp from Sulawesi
Sulawesi ( ), also known as Celebes ( ), is an island in Indonesia. One of the four Greater Sunda Isla ...
'' Cai, Wowor & Choy, 2009
*''
Caridina wumingensis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
Caridina yilong
''Caridina yilong'' is a species of freshwater shrimp in the family Atyidae. ''C. yilong'' was described from Lake Yilong (Yilong Hu) in Yunnan, China, and was believed to be endemic to the margins of the lake among fringing vegetation. Only the ...
phylogenetic
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
studies have questioned the
monophyly
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria:
# the grouping contains its own most recent comm ...
of ''Caridina''.
Threats and Conservation
As of March 2023, the IUCN Red List lists 56 ''Caridina'' species as threatened, with 18 listed as critically endangered, 5 listed as endangered, and 33 listed as vulnerable.
Of these, two (''
Caridina apodosis
''Caridina'' is a genus of freshwater atyid shrimp. They are widely found in tropical or subtropical water in Asia, Oceania and Africa. They are filter-feeders and omnivorous scavengers. They range from 0.9 to 9.8 mm ('' C. cantonensis'') ...
'' and ''
Caridina yilong
''Caridina yilong'' is a species of freshwater shrimp in the family Atyidae. ''C. yilong'' was described from Lake Yilong (Yilong Hu) in Yunnan, China, and was believed to be endemic to the margins of the lake among fringing vegetation. Only the ...
'') are listed as possibly extinct and one ('' Caridina dennerli'') is listed as possibly extinct in the wild.