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''Podarcis carbonelli'', commonly known as Carbonell's wall lizard, is a species of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
in the family
Lacertidae The Lacertidae are the family (biology), family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at least 300 species in 39 genera. They represent the dominant group o ...
. The species is native to Portugal and Spain. This lizard reaches a total length (including tail) of 20 cm (8 in), and feeds primarily on small invertebrates such as insects, arachnids, and snails. Its natural habitats are temperate
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
s and sandy shores. Habitat loss threatens its survival.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''carbonelli'', is in honor of "J. Carbonell" who is the wife of Pérez-Mellado. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Podarcis carbonelli'', p. 47).


Description

Carbonell's wall lizard grows to a snout-to-vent length of with a tail about twice as long. Females tend to be slightly larger than males in some localities. The dorsal surface is usually grey or brown, but is sometimes green (especially so in males), copiously speckled with rows of dark markings. The flanks may also be somewhat greenish with reticulated, dark markings. The underparts are whitish and there are often small blue spots along the edge of the belly. It differs from all other Iberian Wall Lizards by generally showing greenish flanks but no green backs. Carbonell's wall lizard much resembles the closely related
Bocage's wall lizard Bocage's wall lizard (''Podarcis bocagei'') is a species of lizard in the family Lacertidae. The species is endemic to the Iberian Peninsula. Its natural habitats are temperate forests, temperate shrubland, Mediterranean-type shrubby vegetation, s ...
, but that species tends to have more clearly defined markings and a yellow, orange or pink belly, and shows no green flanks and no blue outer ventral scales.


Distribution and habitat

Carbonell's wall lizard is native to Portugal and Spain. Its geographic range consists of a number of isolated populations in western and central Portugal, another in Coto Doñana in southwestern Spain and a separate subspecies is present on the Berlenga Islands off the coast of Portugal. Being specialized on Mediterranean climates with cool to warm summers, it has stable populations at the Atlantic coast, occupying
sand dunes A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...
with barely any vegetation. However, the remaining inland populations in the western mountainous Sistema Central, where it inhabits up to , seem to rapidly decrease, probably a result of global warming.


Behaviour

Carbonell's wall lizard is often seen on dry banks where it may be present in large numbers. It takes refuge in cracks and among tree roots.


Diet

''P. carbonelli'' feeds mainly on arthropods but, particularly on the Berlenga Islands, also consumes snails.


Reproduction

''P. carbonelli'' is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
. In central Portugal there is usually one
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
of two eggs each year, but in the Berlengas, several clutches of up to four eggs are laid. These take ten to fifteen weeks to hatch.


Status

Carbonell's wall lizard has a number of isolated populations and its total geographic range is less than . It lives in oak woodland and
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
and although some populations are in protected areas, others are subject to habitat degradation. Although it is common in some suitable habitats, in general the population is thought to be declining and the International Union for Conservation of Nature has assessed it as being an "
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
".


References


Further reading

*Pérez-Mellado, Valentin (1981). "''Nuevos datos sobre la sistemática y distribucion de'' Podarcis bocagei ''(Seoane, 1884) (Sauria, Lacertidae) en la Península Ibérica'' ". ''Amphibia-Reptilia'' 2: 259-265. (''Podarcis bocagei carbonelli'', new subspecies). (in Spanish). *Sá-Sousa, Paulo; Harris, D. James (2002). "''Podarcis carbonelli'' Pérez-Mellado, 1981 is a distinct species". ''Amphibia-Reptilia'' 23 (4): 459-468.


See Also

The Italian wall lizard, a related species.{{Taxonbar, from=Q2703268 carbonelli Reptiles described in 1981 Lizards of Europe Endemic reptiles of the Iberian Peninsula Taxa named by Valentín Pérez-Mellado Taxonomy articles created by Polbot