Caracanthinae
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''Caracanthus'', the coral crouchers, or orbicular velvetfishes, are a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
es. They live in coral reefs of the tropical Indo-Pacific. This genus is the only member of the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Caracanthinae, part of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Scorpaenidae The Scorpaenidae (also known as scorpionfish) are a family (biology), family of mostly ocean, marine fish that includes many of the world's most venomous species. As their name suggests, scorpionfish have a type of "sting" in the form of sharp ...
.


Taxonomy

''Caracanthus'' was first formally described as a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus in 1845 by the Danish
zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
Henrik Nikolai Krøyer when he described the new species the Hawaiian orbicular velvetfish ('' Caracanthus typicus''). The genus is the only genus in the monotypic subfamily Caracanthinae within the family Scorpaenidae. Molecular studies have found that the Caracanthinae should probably be treated as a tribe within the subfamily
Scorpaeninae Scorpaeninae is a subfamily of ray-finned fish belonging to the family Scorpaenidae in the order Scorpaeniformes, it includes the scorpionfishes, the lionfishes and turkeyfishes. They bear venomous spines in the anal, dorsal and pelvic fins which ...
, or possibly included in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Scorpaenini Scorpaenini is a tribe of marine ray-finned fishes, one of two tribes in the subfamily Scorpaeninae. This tribe contains the "typical" or "true" scorpionfishes. The taxonomy of the scorpionfishes is in some flux, the 5th Edition of Fishes of the ...
. The genus name is a compound of ''cara'', meaning "head" and ''acanthus'', meaning "thorn" or "spine", an allusion to the strong spines on the infraorbital bone of ''C. typicus''.


Characteristics

''Caracanthus'' fishes have an oval, laterally compressed body which is covered in small, rough papillae with a small, terminal mouth. There is a single notch in the dorsal fin which has its origin on the nape and which contains between 6 and 8 spines and 11 and 14 soft rays. The
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
has 2 spines and between 11 and 14 soft rays. There are 12 to 14 rays in the
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
s but the pelvic fins are small and barely noticeable, they have a single spine and 2 or 3 small soft rays. There are tiny scales on the head, each bearing a single spine. Like the related velvetfishes, they have a velvety skin. The small
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
s are used to wedge themselves into crevices in the coral. These small fishes vary in size from a maximum
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
of in ''C. typicus'' to a maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of for the other 3 species.


Species

There are currently four recognized species in this genus: * '' Caracanthus maculatus'' ( J. E. Gray, 1831) (Spotted coral croucher) * '' Caracanthus madagascariensis'' ( Guichenot, 1869) (Spotted croucher) * '' Caracanthus typicus'' Krøyer, 1845 (Hawaiian orbicular velvetfish) * '' Caracanthus unipinna'' ( J. E. Gray, 1831) (Pygmy coral croucher)


Distribution and habitat

''Caracanthus'' fishes are found in the Indian and Pacific oceans where they are closely associated with coral, living among the coral branches.


References

{{Taxonbar, from=Q1592961 Caracanthinae Marine fish genera Taxa named by Henrik Nikolai Krøyer