The New World barbets are a

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

, Capitonidae, of 15 birds in the order

Piciformes Nine families of largely arboreal birds make up the order Piciformes (), the best-known of them being the Picidae, which includes the woodpeckers and close relatives. The Piciformes contain about 71 living genera with a little over 450 species, ...

, which inhabit humid forests in Central and South America. They are closely related to the

toucan Toucans (, ) are Neotropical birds in the family Ramphastidae. They are most closely related to the Semnornis, Toucan barbets. They are brightly marked and have large, often colorful Beak, bills. The family includes five genus, genera and over ...

s. The New World barbets are plump birds, with short necks and large heads. They get their name from the bristles that fringe their heavy bills. Most

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

are brightly coloured and live in tropical forest. These barbets are mostly arboreal birds, which nest in tree holes dug by breeding pairs, laying two to four eggs. They eat fruit and insects. These birds do not

migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...

Taxonomy

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

New World barbets have been found dating from the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...

. The closest relatives of the barbets are the toucans, and these two families are also closely related to the honeyguides and woodpeckers (with which they form the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Piciformes). Formerly, the barbets have been treated as one

. This has turned out to be

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

, though, with regard to toucans; thus, only the New World true barbets are retained in the Capitonidae. The African barbets (

Lybiidae Lybiidae is a family (biology), family of birds also known as the African barbets. There are 44 species ranging from the type genus ''Lybius'' of forest interior to the tinkerbirds (''Pogoniulus'') of forest and scrubland. They are found througho ...

) and the Asian barbets (

Megalaimidae Megalaimidae, the Asian barbets, are a family of birds, comprising two genera with 35 species native to the forests of the Indomalayan realm from Tibet to Indonesia. They were once clubbed with all barbets in the family Capitonidae but the Old Wo ...

), as well as the two toucan-barbets from the Americas (

Semnornithidae The toucan-barbets are small birds in the genus ''Semnornis''. The genus was often included in the paraphyletic barbets, but recently is usually classified into a distinct family, Semnornithidae; alternatively, all barbets might be moved to the ...

) are currently split from this family. Alternatively, the toucans, which

evolved Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...

from a common ancestor shared with the American barbets, might be included in the traditional all-encompassing barbet family. As they have evolved characteristics that are unique to themselves, they are usually treated separately, thus the barbets are split up according to the four lineages. The

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

relationship between the New World barbets and the eight other families in the

is shown in the cladogram below. The number of species in each family is taken from the list maintained by Frank Gill, Pamela C. Rasmussen and David Donsker on behalf of the

International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...

(IOC).

Fossils

* Genus '' Capitonides'' (Early – Middle Miocene of Europe) (fossil) **†''Capitonides europeus''

Ecology

While most New World barbet species inhabit lowland forest, some range into montane and temperate forests, as well. Most are restricted to habitats containing trees with dead wood, which are used for nesting. The diet of barbets is mixed, with fruit being the dominant part. Small prey items are also taken, especially when nesting. Barbets are capable of shifting their diet quickly in the face of changing food availability. Numerous species of fruiting trees and bushes are visited; an individual barbet may feed on as many as 60 different species in its range. They also visit plantations and take cultivated fruit and vegetables. Fruit is eaten whole, and indigestible material such as

seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...

pits is regurgitated later (often before

singing Singing is the art of creating music with the voice. It is the oldest form of musical expression, and the human voice can be considered the first musical instrument. The definition of singing varies across sources. Some sources define singi ...

). Regurgitation does not usually happen in the nest (as happens with

s). Like their relatives, New World barbets are thought to be important agents in

seed dispersal In spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...

in tropical forests. As well as taking fruit, they also take

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

prey, gleaned from the branches and trunks of trees. A wide range of insects is taken, including ants, beetles, and moths. Scorpions and centipedes are also taken, and a few species take small vertebrates such as frogs.

Relationship with humans

New World barbets have little direct impact on humans. The loss of forest can have a deleterious effect on barbet species dependent on old growth, to the benefit of species that favor more disturbed or open habitat. Three species of New World barbets are listed as threatened by the

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

: The white-mantled barbet of

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

is listed as

endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...

and the

five-coloured barbet The five-colored barbet (''Capito quinticolor'') is a species of bird in the family Capitonidae, the New World barbets. It is found in Colombia and Ecuador. Taxonomy and systematics The five-colored barbet is monotypic. It and the white-mant ...

as vulnerable, the two having a relatively small range threatened by

deforestation Deforestation or forest clearance is the removal and destruction of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. Ab ...

for the

timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...

industry and to create space for agriculture (including

coca Coca is any of the four cultivated plants in the family Erythroxylaceae, native to western South America. Coca is known worldwide for its psychoactive alkaloid, cocaine. Coca leaves contain cocaine which acts as a mild stimulant when chewed or ...

and

marijuana Cannabis (), commonly known as marijuana (), weed, pot, and ganja, List of slang names for cannabis, among other names, is a non-chemically uniform psychoactive drug from the ''Cannabis'' plant. Native to Central or South Asia, cannabis has ...

) and livestock, and mining. The quite recently discovered scarlet-banded barbet of

Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...

is considered vulnerable due to its small population size (estimated at under 1000 birds), although its remote habitat is not immediately threatened.

References

External links

Barbet videos
on the Internet Bird Collection

{{Taxonbar, from=Q809613 Bird families Birds of South America Birds of the Americas Taxa named by Charles Lucien Bonaparte