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''Canthium'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules ...
. They are shrubs and small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s. The leaves are
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...
and the stems are usually thorny.


Distribution

''Canthium'' species are predominantly found in
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, especially in
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
and the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
. A small number of species is found in
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, and
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world, with a population exceeding 165 million pe ...
. Only a limited number of species is found on the African continent, especially in
Southern Southern may refer to: Businesses * China Southern Airlines, airline based in Guangzhou, China * Southern Airways, defunct US airline * Southern Air, air cargo transportation company based in Norwalk, Connecticut, US * Southern Airways Express, M ...
and
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the histori ...
.


Taxonomy

''Canthium'' was named by
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biolog ...
in 1785 in
Encyclopédie Méthodique The ''Encyclopédie méthodique par ordre des matières'' ("Methodical Encyclopedia by Order of Subject Matter") was published between 1782 and 1832 by the French publisher Charles Joseph Panckoucke, his son-in-law Henri Agasse, and the latter's ...
. The name is a latinisation of "kantankara", a
Malayalam Malayalam (; , ) is a Dravidian language spoken in the Indian state of Kerala and the union territories of Lakshadweep and Puducherry ( Mahé district) by the Malayali people. It is one of 22 scheduled languages of India. Malayalam wa ...
name from
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South C ...
for ''Canthium coromandelicum''. ''Kantan'' means "shining" and ''kara'' means "a spiny shrub". The
biological type In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...
for the genus consists of
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository ...
s originally described by
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biolog ...
as ''Canthium parviflorum''''Canthium'' In: Index Nominum Genericorum. In:
Regnum Vegetabile The International Association for Plant Taxonomy (IAPT) is an organization established to promote an understanding of plant biodiversity, facilitate international communication of research between botanists, and oversee matters of uniformity and ...
but this species is now included in ''
Canthium coromandelicum ''Canthium coromandelicum'' is a bushy thorny suffruticose herb, a native of India found mainly in the Coromandel region. Description ''Canthium coromandelicum'' is a shrub, usually with opposite horizontal thorns a little above the leaf. Some ...
''. ''Canthium'' is a member of Vanguerieae, a
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...
that is
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
and easily recognized morphologically, but in which
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
boundaries were, for a long time, very unclear. ''Canthium'' was especially problematic, and until the 1980s, it was defined broadly and known to be
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
. ''
Psydrax ''Psydrax'' is a genus of flowering plants in the family Rubiaceae. It consists of trees, shrubs, and a few lianas in the paleotropics. Taxonomy The genus was named by Joseph Gaertner in 1788 in his book, '' De Fructibus et Seminibus Plantaru ...
'' was separated from it in 1985, as was '' Keetia'' in 1986. These were followed by ''
Pyrostria ''Pyrostria'' is a genus of dioecious flowering plants in the Family (biology), family Rubiaceae. Most of the species are endemic to Madagascar, others occur on islands in the western Indian Ocean (Mauritius, Comoros, Réunion, Rodrigues, Socotr ...
'' and ''
Multidentia ''Multidentia'' is a genus of flowering plants in the Family (biology), family Rubiaceae. Distribution ''Multidentia'' is restricted to tropical Africa with the greatest concentration of species occurring in eastern Africa. Taxonomy It was orig ...
'' in 1987. The
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed betw ...
''
Afrocanthium ''Afrocanthium'' is a genus of flowering plants in the family Rubiaceae. It consists of deciduous, unarmed trees, and shrubs. They are native to East Africa, from Sudan and Ethiopia to South Africa. Taxonomy ''Afrocanthium'' was first recogniz ...
'' was raised to generic
rank Rank is the relative position, value, worth, complexity, power, importance, authority, level, etc. of a person or object within a ranking, such as: Level or position in a hierarchical organization * Academic rank * Diplomatic rank * Hierarchy * H ...
in 2004, followed by '' Bullockia'' in 2009. A few species were transferred to ''Canthium'' from '' Rytigynia'' and other genera in 2004. The genus was further reduced by the transfer of species to '' Peponidium'' and ''
Pyrostria ''Pyrostria'' is a genus of dioecious flowering plants in the Family (biology), family Rubiaceae. Most of the species are endemic to Madagascar, others occur on islands in the western Indian Ocean (Mauritius, Comoros, Réunion, Rodrigues, Socotr ...
''. In 2016, two ''Canthium'' species endemic to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
were transferred to a genus of their own, ''
Kanapia ''Kanapia'' is a genus of flowering plants in the family Rubiaceae. The genus is endemic to the Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no ...
''. The final circumscription of ''Canthium'' will remain in doubt until
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
studies achieve greater
resolution Resolution(s) may refer to: Common meanings * Resolution (debate), the statement which is debated in policy debate * Resolution (law), a written motion adopted by a deliberative body * New Year's resolution, a commitment that an individual ma ...
for the clade containing ''
Canthium coromandelicum ''Canthium coromandelicum'' is a bushy thorny suffruticose herb, a native of India found mainly in the Coromandel region. Description ''Canthium coromandelicum'' is a shrub, usually with opposite horizontal thorns a little above the leaf. Some ...
'' and its closest relatives.


Species

,
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants ...
recognises the following species: * ''
Canthium aciculatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
* '' Canthium angustifolium''
Roxb. William Roxburgh FRSE FRCPE FLS (3/29 June 1751 – 18 February 1815) was a Scottish surgeon and botanist who worked extensively in India, describing species and working on economic botany. He is known as the founding father of Indian botany. ...
* ''
Canthium arboreum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' Vidal * ''
Canthium aurantiacum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* '' Canthium berberidifolium'' Geddes * ''
Canthium bipinnatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
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)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium brunneum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium calvum'' Craib * '' Canthium cambodianum'' Pit. * ''
Canthium campanulatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' Thwaites * ''
Canthium cavaleriei ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' H.Lév. * ''
Canthium ciliatum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' (D.Dietr.)
Kuntze Kuntze is a surname of German origin. People with that name include: * Carl Kuntze (1922-2006), Dutch rower who competed at the 1952 Summer Olympics * Edward J. Kuntze (1826-1870), Prussian-born American sculptor * Otto Kuntze Carl Ernst Otto K ...
* ''
Canthium coffeoides ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Pierre ex Pit. * ''
Canthium congestiflorum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
* ''
Canthium cordatum ''Canthium'' is a genus of flowering plants in the Family (biology), family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually Thorns, spines, and prickles, thorny. Distribution ''Canthium'' species a ...
'' Dillwyn * ''
Canthium coromandelicum ''Canthium coromandelicum'' is a bushy thorny suffruticose herb, a native of India found mainly in the Coromandel region. Description ''Canthium coromandelicum'' is a shrub, usually with opposite horizontal thorns a little above the leaf. Some ...
'' ( Brum.f.) Alston * '' Canthium depressinerve''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
* ''
Canthium ellipticum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium fenicis'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium ferrugineum'' Craib * '' Canthium filipendulum'' Pierre ex Pit. * '' Canthium fraternum''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium glaucum''
Hiern Hiern is a British surname. Notable people with the surname include: * Barry Hiern (born 1951), Australian cricketer *Ross Hiern (1922–1999), Australian cricketer *William Philip Hiern (1839–1925), British mathematician and botanist See also * ...
* ''
Canthium gracilipes ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Kurz Sebastian Kurz (; born 27 August 1986) is a former Austrian politician who twice served as chancellor of Austria, initially from December 2017 to May 2019 and then a second time from January 2020 to October 2021. Kurz was born and raised in ...
* ''
Canthium hainanense ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
) Lantz
* ''
Canthium hirtellum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
* ''
Canthium hispidonervosum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' ( De Wild.) C.M.Evrard * '' Canthium horridulum'' Craib * ''
Canthium horridum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Blume * '' Canthium inerme'' ( L.f.)
Kuntze Kuntze is a surname of German origin. People with that name include: * Carl Kuntze (1922-2006), Dutch rower who competed at the 1952 Summer Olympics * Edward J. Kuntze (1826-1870), Prussian-born American sculptor * Otto Kuntze Carl Ernst Otto K ...
* ''
Canthium korthalsianum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium kuntzeanum'' Bridson * '' Canthium laeve'' Teijsm. & Binn. * ''
Canthium lasianthoides ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium libericum'' Dinkl. * ''
Canthium longipes ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Geddes * ''
Canthium lucidum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' R.Br. * '' Canthium macrocarpum'' Thwaites * '' Canthium malayense'' K.M.Wong * ''
Canthium megacarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium megistocarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* '' Canthium merrillianum''
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* '' Canthium merrillii'' (Setch.)
Christoph. Erling Christophersen (April 17, 1898 – November 9, 1994) was a Norwegian botanist, geographer and diplomat. He participated in and led several notable scientific expeditions in the 20th century, including the fifth Tanager Expedition (1924) to ...
* ''
Canthium mite ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Bartl. Friedrich Gottlieb Bartling (December 9, 1798 – November 20, 1875) was a German botanist who was a native of Hanover. He studied natural sciences at the University of Göttingen, and in 1818 took a botanical journey through Hungary and Croatia ...
ex
DC. Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...
* '' Canthium molle''
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Gamble Gambling (also known as betting or gaming) is the wagering of something of value ("the stakes") on a random event with the intent of winning something else of value, where instances of strategy are discounted. Gambling thus requires three ele ...
* ''
Canthium moluccanum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Roxb. William Roxburgh FRSE FRCPE FLS (3/29 June 1751 – 18 February 1815) was a Scottish surgeon and botanist who worked extensively in India, describing species and working on economic botany. He is known as the founding father of Indian botany. ...
* '' Canthium oblongum'' (Valeton) Kaneh. * ''
Canthium oliganthum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
) Boerl.
* ''
Canthium oligocarpum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
''
Hiern Hiern is a British surname. Notable people with the surname include: * Barry Hiern (born 1951), Australian cricketer *Ross Hiern (1922–1999), Australian cricketer *William Philip Hiern (1839–1925), British mathematician and botanist See also * ...
* ''
Canthium parvifolium ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
Roxb. William Roxburgh FRSE FRCPE FLS (3/29 June 1751 – 18 February 1815) was a Scottish surgeon and botanist who worked extensively in India, describing species and working on economic botany. He is known as the founding father of Indian botany. ...
* ''
Canthium paucinervium ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium pedunculare ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Cav. * ''
Canthium perakanthus ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' ined. * '' Canthium polyanthum''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium puberulum'' Thwaites ex Hook.f. * ''
Canthium quadratum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' Craib * ''
Canthium rheedei ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
''
DC. Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...
* '' Canthium sarcocarpum'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium sarmentosum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' Craib * '' Canthium scabridum''
Ridl. Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
* ''
Canthium scandens ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' Blume * ''
Canthium schlechterianum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& L.M.Perry
* ''
Canthium simile ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
''
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
& Chun
* ''
Canthium sordidum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
'' (
K.Schum. Karl Moritz Schumann (17 June 1851 – 22 March 1904) was a German botanist. Schumann was born in Görlitz. He was curator of the Botanisches Museum in Berlin-Dahlem from 1880 until 1894. He also served as the first chairman of the ''Deutsc ...
) Bullock
* ''
Canthium spirostylum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium stellulatum'' Craib * '' Canthium strigosum'' Craib * '' Canthium strychnoides'' Craib * ''
Canthium subaureum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' Craib * '' Canthium subcapitatum'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* '' Canthium suberosum'' Codd * ''
Canthium subsessilifolium ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia ...
'' (
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium tavoyanum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' (R.Parker)
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium travancoricum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
''
Bedd. Colonel Richard Henry Beddome (11 May 1830 – 23 February 1911) was a British military officer and Natural history, naturalist in India, who became chief conservator of the Tamil Nadu Forest Department, Madras Forest Department. In the mid- ...
* '' Canthium trichophorum'' Quisumb. &
Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...
* ''
Canthium umbelligerum ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espe ...
''
Miq. Friedrich Anton Wilhelm Miquel (24 October 1811 – 23 January 1871) was a Dutch botanist, whose main focus of study was on the flora of the Dutch East Indies. Early life Miquel was born in Neuenhaus and studied medicine at the University of Gro ...
* '' Canthium vanwykii'' Tilney & Kok * ''
Canthium villarii ''Canthium'' is a genus of flowering plants in the family Rubiaceae. They are shrubs and small trees. The leaves are deciduous and the stems are usually thorny. Distribution ''Canthium'' species are predominantly found in Southeast Asia, espec ...
'' Vidal * '' Canthium violaceum'' Zoll. & Moritzi


References

{{Taxonbar, from=Q3244344 Rubiaceae genera Afrotropical realm flora Indomalayan realm flora Taxa named by Jean-Baptiste Lamarck Taxonomy articles created by Polbot